NATURE . COM {}

Matching Content Categories {📚}

• Education
• Science
• Health & Fitness

Content Management System {📝}

What CMS is nature.com built with?

Custom-built

No common CMS systems were detected on Nature.com, and no known web development framework was identified.

Traffic Estimate {📈}

What is the average monthly size of nature.com audience?

🌠 Phenomenal Traffic: 5M - 10M visitors per month


Based on our best estimate, this website will receive around 5,000,019 visitors per month in the current month.
However, some sources were not loaded, we suggest to reload the page to get complete results.

check SE Ranking
check Ahrefs
check Similarweb
check Ubersuggest
check Semrush

How Does Nature.com Make Money? {💸}

Display Ads {🎯}

Direct Advertisers (10)

Reseller Advertisers (38)

How Much Does Nature.com Make? {💰}

Display Ads {🎯}

$63,100 per month
Estimations show Nature.com's display ad online revenue falls between $42,042 and $115,616 per month.

Keywords {🔍}

article, arthritis, rheumatoid, google, scholar, nature, cas, loci, genet, association, research, university, genetics, health, nat, medical, data, paul, access, department, identifies, study, genomewide, usa, institute, manchester, supplementary, content, genetic, eyre, bakker, wellcome, trust, raychaudhuri, national, cookies, analysis, diogo, stahl, trynka, authors, snp, genotyping, studies, metaanalysis, risk, gene, hospital, center, robert,

Topics {✒️}

nature portfolio permissions reprints privacy policy advertising nature 434 nature social media identify rheumatoid-arthritis–susceptibility loci swedish research council genome-wide association studies genomics study syndicate x-linked gene expression suna onengut-gumuscu & stephen health research population-based linkage analyses musculoskeletal research rna-seq data medical research author correspondence harm-jan westra chiea chuen khor instituto de fundación imputation-driven meta-analysis springerlink instant access mendelian randomisation analysis snp&seq technology platform permissions public health genomics meta-analysis identifies genome-wide identification rheumatoid arthritis genetics plenge & soumya raychaudhuri seropositive rheumatoid arthritis rheumatoid arthritis regions burroughs wellcome fund dense genotyping identifies privacy immunological genome project european economic area bayesian inference analyses independent genetic variants harvard medical school national institutes health competing financial interests collaborative meta-analysis arthritis foundation personal data explore content subscription content swedish umea samples

Schema {🗺️}

WebPage:
      mainEntity:
         headline:High-density genetic mapping identifies new susceptibility loci for rheumatoid arthritis
         description:Jane Worthington and colleagues use a high-density genotyping chip to identify new susceptibility loci for rheumatoid arthritis and examine genetic overlap with other autoimmune diseases. Their results increase the number of confirmed rheumatoid arthritis risk loci in individuals of European ancestry and refine the location of many previously identified association signals to single genes. Using the Immunochip custom SNP array, which was designed for dense genotyping of 186 loci identified through genome-wide association studies (GWAS), we analyzed 11,475 individuals with rheumatoid arthritis (cases) of European ancestry and 15,870 controls for 129,464 markers. We combined these data in a meta-analysis with GWAS data from additional independent cases (n = 2,363) and controls (n = 17,872). We identified 14 new susceptibility loci, 9 of which were associated with rheumatoid arthritis overall and five of which were specifically associated with disease that was positive for anticitrullinated peptide antibodies, bringing the number of confirmed rheumatoid arthritis risk loci in individuals of European ancestry to 46. We refined the peak of association to a single gene for 19 loci, identified secondary independent effects at 6 loci and identified association to low-frequency variants at 4 loci. Bioinformatic analyses generated strong hypotheses for the causal SNP at seven loci. This study illustrates the advantages of dense SNP mapping analysis to inform subsequent functional investigations.
         datePublished:2012-11-11T00:00:00Z
         dateModified:2012-11-11T00:00:00Z
         pageStart:1336
         pageEnd:1340
         sameAs:https://doi.org/10.1038/ng.2462
         keywords:
            Genetic association study
            Rheumatoid arthritis
            Biomedicine
            general
            Human Genetics
            Cancer Research
            Agriculture
            Gene Function
            Animal Genetics and Genomics
         image:
            https://media.springernature.com/lw1200/springer-static/image/art%3A10.1038%2Fng.2462/MediaObjects/41588_2012_Article_BFng2462_Fig1_HTML.jpg
         isPartOf:
            name:Nature Genetics
            issn:
               1546-1718
               1061-4036
            volumeNumber:44
            type:
               Periodical
               PublicationVolume
         publisher:
            name:Nature Publishing Group US
            logo:
               url:https://www.springernature.com/app-sn/public/images/logo-springernature.png
               type:ImageObject
            type:Organization
         author:
               name:Steve Eyre
               affiliation:
                     name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre
                     address:
                        name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre, Manchester, UK
                        type:PostalAddress
                     type:Organization
                     name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre
                     address:
                        name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre, Manchester, UK
                        type:PostalAddress
                     type:Organization
               type:Person
               name:John Bowes
               affiliation:
                     name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre
                     address:
                        name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre, Manchester, UK
                        type:PostalAddress
                     type:Organization
                     name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre
                     address:
                        name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre, Manchester, UK
                        type:PostalAddress
                     type:Organization
               type:Person
               name:Dorothée Diogo
               affiliation:
                     name:Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School
                     address:
                        name:Division of Rheumatology, Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
                        type:PostalAddress
                     type:Organization
                     name:Brigham and Women's Hospital, Harvard Medical School
                     address:
                        name:Division of Genetics, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
                        type:PostalAddress
                     type:Organization
                     name:Program in Medical and Population Genetics, Broad Institute
                     address:
                        name:Program in Medical and Population Genetics, Broad Institute, Cambridge, USA
                        type:PostalAddress
                     type:Organization
               type:Person
               name:Annette Lee
               affiliation:
                     name:The Feinstein Institute for Medical Research, North Shore–Long Island Jewish Health System
                     address:
                        name:The Feinstein Institute for Medical Research, North Shore–Long Island Jewish Health System, Manhasset, USA
                        type:PostalAddress
                     type:Organization
               type:Person
               name:Anne Barton
               affiliation:
                     name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre
                     address:
                        name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre, Manchester, UK
                        type:PostalAddress
                     type:Organization
                     name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre
                     address:
                        name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre, Manchester, UK
                        type:PostalAddress
                     type:Organization
               type:Person
               name:Paul Martin
               affiliation:
                     name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre
                     address:
                        name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre, Manchester, UK
                        type:PostalAddress
                     type:Organization
                     name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre
                     address:
                        name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre, Manchester, UK
                        type:PostalAddress
                     type:Organization
               type:Person
               name:Alexandra Zhernakova
               affiliation:
                     name:Leiden University Medical Centre
                     address:
                        name:Department of Rheumatology, Leiden University Medical Centre, Leiden, The Netherlands
                        type:PostalAddress
                     type:Organization
                     name:University Medical Center Groningen and University of Groningen
                     address:
                        name:Department of Genetics, University Medical Center Groningen and University of Groningen, Groningen, The Netherlands
                        type:PostalAddress
                     type:Organization
               type:Person
               name:Eli Stahl
               affiliation:
                     name:Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School
                     address:
                        name:Division of Rheumatology, Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
                        type:PostalAddress
                     type:Organization
                     name:Brigham and Women's Hospital, Harvard Medical School
                     address:
                        name:Division of Genetics, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
                        type:PostalAddress
                     type:Organization
                     name:Program in Medical and Population Genetics, Broad Institute
                     address:
                        name:Program in Medical and Population Genetics, Broad Institute, Cambridge, USA
                        type:PostalAddress
                     type:Organization
               type:Person
               name:Sebastien Viatte
               affiliation:
                     name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre
                     address:
                        name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre, Manchester, UK
                        type:PostalAddress
                     type:Organization
                     name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre
                     address:
                        name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre, Manchester, UK
                        type:PostalAddress
                     type:Organization
               type:Person
               name:Kate McAllister
               affiliation:
                     name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre
                     address:
                        name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre, Manchester, UK
                        type:PostalAddress
                     type:Organization
                     name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre
                     address:
                        name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre, Manchester, UK
                        type:PostalAddress
                     type:Organization
               type:Person
               name:Christopher I Amos
               affiliation:
                     name:University of Texas MD Anderson Cancer Center
                     address:
                        name:University of Texas MD Anderson Cancer Center, Houston, USA
                        type:PostalAddress
                     type:Organization
               type:Person
               name:Leonid Padyukov
               affiliation:
                     name:Rheumatology Unit, Karolinska Institutet and Karolinska University Hospital Solna
                     address:
                        name:Department of Medicine, Rheumatology Unit, Karolinska Institutet and Karolinska University Hospital Solna, Stockholm, Sweden
                        type:PostalAddress
                     type:Organization
               type:Person
               name:Rene E M Toes
               affiliation:
                     name:Leiden University Medical Centre
                     address:
                        name:Department of Rheumatology, Leiden University Medical Centre, Leiden, The Netherlands
                        type:PostalAddress
                     type:Organization
               type:Person
               name:Tom W J Huizinga
               affiliation:
                     name:Leiden University Medical Centre
                     address:
                        name:Department of Rheumatology, Leiden University Medical Centre, Leiden, The Netherlands
                        type:PostalAddress
                     type:Organization
               type:Person
               name:Cisca Wijmenga
               affiliation:
                     name:University Medical Center Groningen and University of Groningen
                     address:
                        name:Department of Genetics, University Medical Center Groningen and University of Groningen, Groningen, The Netherlands
                        type:PostalAddress
                     type:Organization
               type:Person
               name:Gosia Trynka
               affiliation:
                     name:Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School
                     address:
                        name:Division of Rheumatology, Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
                        type:PostalAddress
                     type:Organization
                     name:Brigham and Women's Hospital, Harvard Medical School
                     address:
                        name:Division of Genetics, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
                        type:PostalAddress
                     type:Organization
                     name:Program in Medical and Population Genetics, Broad Institute
                     address:
                        name:Program in Medical and Population Genetics, Broad Institute, Cambridge, USA
                        type:PostalAddress
                     type:Organization
                     name:University Medical Center Groningen and University of Groningen
                     address:
                        name:Department of Genetics, University Medical Center Groningen and University of Groningen, Groningen, The Netherlands
                        type:PostalAddress
                     type:Organization
               type:Person
               name:Lude Franke
               affiliation:
                     name:University Medical Center Groningen and University of Groningen
                     address:
                        name:Department of Genetics, University Medical Center Groningen and University of Groningen, Groningen, The Netherlands
                        type:PostalAddress
                     type:Organization
               type:Person
               name:Harm-Jan Westra
               affiliation:
                     name:University Medical Center Groningen and University of Groningen
                     address:
                        name:Department of Genetics, University Medical Center Groningen and University of Groningen, Groningen, The Netherlands
                        type:PostalAddress
                     type:Organization
               type:Person
               name:Lars Alfredsson
               affiliation:
                     name:Karolinska Institutet
                     address:
                        name:Department of Environmental Medicine, Karolinska Institutet, Stockholm, Sweden
                        type:PostalAddress
                     type:Organization
               type:Person
               name:Xinli Hu
               affiliation:
                     name:Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School
                     address:
                        name:Division of Rheumatology, Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
                        type:PostalAddress
                     type:Organization
                     name:Brigham and Women's Hospital, Harvard Medical School
                     address:
                        name:Division of Genetics, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
                        type:PostalAddress
                     type:Organization
                     name:Program in Medical and Population Genetics, Broad Institute
                     address:
                        name:Program in Medical and Population Genetics, Broad Institute, Cambridge, USA
                        type:PostalAddress
                     type:Organization
                     name:Harvard–Massachusetts Institute of Technology Division of Health Sciences and Technology
                     address:
                        name:Harvard–Massachusetts Institute of Technology Division of Health Sciences and Technology, Boston, USA
                        type:PostalAddress
                     type:Organization
               type:Person
               name:Cynthia Sandor
               affiliation:
                     name:Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School
                     address:
                        name:Division of Rheumatology, Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
                        type:PostalAddress
                     type:Organization
                     name:Brigham and Women's Hospital, Harvard Medical School
                     address:
                        name:Division of Genetics, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
                        type:PostalAddress
                     type:Organization
                     name:Program in Medical and Population Genetics, Broad Institute
                     address:
                        name:Program in Medical and Population Genetics, Broad Institute, Cambridge, USA
                        type:PostalAddress
                     type:Organization
               type:Person
               name:Paul I W de Bakker
               affiliation:
                     name:Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School
                     address:
                        name:Division of Rheumatology, Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
                        type:PostalAddress
                     type:Organization
                     name:Brigham and Women's Hospital, Harvard Medical School
                     address:
                        name:Division of Genetics, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
                        type:PostalAddress
                     type:Organization
                     name:Program in Medical and Population Genetics, Broad Institute
                     address:
                        name:Program in Medical and Population Genetics, Broad Institute, Cambridge, USA
                        type:PostalAddress
                     type:Organization
                     name:University Medical Center Utrecht
                     address:
                        name:Department of Epidemiology, University Medical Center Utrecht, Utrecht, The Netherlands
                        type:PostalAddress
                     type:Organization
                     name:University Medical Center Utrecht
                     address:
                        name:Department of Medical Genetics, University Medical Center Utrecht, Utrecht, The Netherlands
                        type:PostalAddress
                     type:Organization
               type:Person
               name:Sonia Davila
               affiliation:
                     name:Genome Institute of Singapore
                     address:
                        name:Division of Human Genetics, Genome Institute of Singapore, Singapore
                        type:PostalAddress
                     type:Organization
               type:Person
               name:Chiea Chuen Khor
               affiliation:
                     name:Genome Institute of Singapore
                     address:
                        name:Division of Human Genetics, Genome Institute of Singapore, Singapore
                        type:PostalAddress
                     type:Organization
               type:Person
               name:Khai Koon Heng
               affiliation:
                     name:Genome Institute of Singapore
                     address:
                        name:Division of Human Genetics, Genome Institute of Singapore, Singapore
                        type:PostalAddress
                     type:Organization
               type:Person
               name:Robert Andrews
               affiliation:
                     name:The Wellcome Trust Sanger Institute
                     address:
                        name:The Wellcome Trust Sanger Institute, Cambridge, UK
                        type:PostalAddress
                     type:Organization
               type:Person
               name:Sarah Edkins
               affiliation:
                     name:The Wellcome Trust Sanger Institute
                     address:
                        name:The Wellcome Trust Sanger Institute, Cambridge, UK
                        type:PostalAddress
                     type:Organization
               type:Person
               name:Sarah E Hunt
               affiliation:
                     name:The Wellcome Trust Sanger Institute
                     address:
                        name:The Wellcome Trust Sanger Institute, Cambridge, UK
                        type:PostalAddress
                     type:Organization
               type:Person
               name:Cordelia Langford
               affiliation:
                     name:The Wellcome Trust Sanger Institute
                     address:
                        name:The Wellcome Trust Sanger Institute, Cambridge, UK
                        type:PostalAddress
                     type:Organization
               type:Person
               name:Deborah Symmons
               affiliation:
                     name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre
                     address:
                        name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre, Manchester, UK
                        type:PostalAddress
                     type:Organization
                     name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre
                     address:
                        name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre, Manchester, UK
                        type:PostalAddress
                     type:Organization
               type:Person
               name:Pat Concannon
               affiliation:
                     name:Center for Public Health Genomics, University of Virginia
                     address:
                        name:Center for Public Health Genomics, University of Virginia, Charlottesville, USA
                        type:PostalAddress
                     type:Organization
               type:Person
               name:Suna Onengut-Gumuscu
               affiliation:
                     name:Center for Public Health Genomics, University of Virginia
                     address:
                        name:Center for Public Health Genomics, University of Virginia, Charlottesville, USA
                        type:PostalAddress
                     type:Organization
               type:Person
               name:Stephen S Rich
               affiliation:
                     name:Center for Public Health Genomics, University of Virginia
                     address:
                        name:Center for Public Health Genomics, University of Virginia, Charlottesville, USA
                        type:PostalAddress
                     type:Organization
               type:Person
               name:Panos Deloukas
               affiliation:
                     name:The Wellcome Trust Sanger Institute
                     address:
                        name:The Wellcome Trust Sanger Institute, Cambridge, UK
                        type:PostalAddress
                     type:Organization
               type:Person
               name:Miguel A Gonzalez-Gay
               affiliation:
                     name:Hospital Marques de Valdecilla, Instituto de Fundación e Investigación Marqués de Valdecilla
                     address:
                        name:Department of Rheumatology, Hospital Marques de Valdecilla, Instituto de Fundación e Investigación Marqués de Valdecilla, Santander, Spain
                        type:PostalAddress
                     type:Organization
               type:Person
               name:Luis Rodriguez-Rodriguez
               affiliation:
                     name:Hospital Clinico San Carlos
                     address:
                        name:Hospital Clinico San Carlos, Madrid, Spain
                        type:PostalAddress
                     type:Organization
               type:Person
               name:Lisbeth Ärlsetig
               affiliation:
                     name:UmeÃ¥ University
                     address:
                        name:Department of Public Health and Clinical Medicine, UmeÃ¥ University, UmeÃ¥, Sweden
                        type:PostalAddress
                     type:Organization
                     name:UmeÃ¥ University
                     address:
                        name:Department of Rheumatology, UmeÃ¥ University, UmeÃ¥, Sweden
                        type:PostalAddress
                     type:Organization
               type:Person
               name:Javier Martin
               affiliation:
                     name:El Instituto de Parasitología y Biomedicina 'López-Neyra'–Consejo Superior de Investigaciones Científicas, Avenida del Conocimiento s/n
                     address:
                        name:El Instituto de Parasitología y Biomedicina 'López-Neyra'–Consejo Superior de Investigaciones Científicas, Avenida del Conocimiento s/n, Granada, Spain
                        type:PostalAddress
                     type:Organization
               type:Person
               name:Solbritt Rantapää-Dahlqvist
               affiliation:
                     name:UmeÃ¥ University
                     address:
                        name:Department of Public Health and Clinical Medicine, UmeÃ¥ University, UmeÃ¥, Sweden
                        type:PostalAddress
                     type:Organization
                     name:UmeÃ¥ University
                     address:
                        name:Department of Rheumatology, UmeÃ¥ University, UmeÃ¥, Sweden
                        type:PostalAddress
                     type:Organization
               type:Person
               name:Robert M Plenge
               affiliation:
                     name:Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School
                     address:
                        name:Division of Rheumatology, Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
                        type:PostalAddress
                     type:Organization
                     name:Brigham and Women's Hospital, Harvard Medical School
                     address:
                        name:Division of Genetics, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
                        type:PostalAddress
                     type:Organization
                     name:Program in Medical and Population Genetics, Broad Institute
                     address:
                        name:Program in Medical and Population Genetics, Broad Institute, Cambridge, USA
                        type:PostalAddress
                     type:Organization
               type:Person
               name:Soumya Raychaudhuri
               affiliation:
                     name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre
                     address:
                        name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre, Manchester, UK
                        type:PostalAddress
                     type:Organization
                     name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre
                     address:
                        name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre, Manchester, UK
                        type:PostalAddress
                     type:Organization
                     name:Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School
                     address:
                        name:Division of Rheumatology, Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
                        type:PostalAddress
                     type:Organization
                     name:Brigham and Women's Hospital, Harvard Medical School
                     address:
                        name:Division of Genetics, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
                        type:PostalAddress
                     type:Organization
                     name:Program in Medical and Population Genetics, Broad Institute
                     address:
                        name:Program in Medical and Population Genetics, Broad Institute, Cambridge, USA
                        type:PostalAddress
                     type:Organization
               type:Person
               name:Lars Klareskog
               affiliation:
                     name:Rheumatology Unit, Karolinska Institutet and Karolinska University Hospital Solna
                     address:
                        name:Department of Medicine, Rheumatology Unit, Karolinska Institutet and Karolinska University Hospital Solna, Stockholm, Sweden
                        type:PostalAddress
                     type:Organization
               type:Person
               name:Peter K Gregersen
               affiliation:
                     name:The Feinstein Institute for Medical Research, North Shore–Long Island Jewish Health System
                     address:
                        name:The Feinstein Institute for Medical Research, North Shore–Long Island Jewish Health System, Manhasset, USA
                        type:PostalAddress
                     type:Organization
               type:Person
               name:Jane Worthington
               affiliation:
                     name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre
                     address:
                        name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre, Manchester, UK
                        type:PostalAddress
                     type:Organization
                     name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre
                     address:
                        name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre, Manchester, UK
                        type:PostalAddress
                     type:Organization
               email:[email protected]
               type:Person
         isAccessibleForFree:
         hasPart:
            isAccessibleForFree:
            cssSelector:.main-content
            type:WebPageElement
         type:ScholarlyArticle
      context:https://schema.org
ScholarlyArticle:
      headline:High-density genetic mapping identifies new susceptibility loci for rheumatoid arthritis
      description:Jane Worthington and colleagues use a high-density genotyping chip to identify new susceptibility loci for rheumatoid arthritis and examine genetic overlap with other autoimmune diseases. Their results increase the number of confirmed rheumatoid arthritis risk loci in individuals of European ancestry and refine the location of many previously identified association signals to single genes. Using the Immunochip custom SNP array, which was designed for dense genotyping of 186 loci identified through genome-wide association studies (GWAS), we analyzed 11,475 individuals with rheumatoid arthritis (cases) of European ancestry and 15,870 controls for 129,464 markers. We combined these data in a meta-analysis with GWAS data from additional independent cases (n = 2,363) and controls (n = 17,872). We identified 14 new susceptibility loci, 9 of which were associated with rheumatoid arthritis overall and five of which were specifically associated with disease that was positive for anticitrullinated peptide antibodies, bringing the number of confirmed rheumatoid arthritis risk loci in individuals of European ancestry to 46. We refined the peak of association to a single gene for 19 loci, identified secondary independent effects at 6 loci and identified association to low-frequency variants at 4 loci. Bioinformatic analyses generated strong hypotheses for the causal SNP at seven loci. This study illustrates the advantages of dense SNP mapping analysis to inform subsequent functional investigations.
      datePublished:2012-11-11T00:00:00Z
      dateModified:2012-11-11T00:00:00Z
      pageStart:1336
      pageEnd:1340
      sameAs:https://doi.org/10.1038/ng.2462
      keywords:
         Genetic association study
         Rheumatoid arthritis
         Biomedicine
         general
         Human Genetics
         Cancer Research
         Agriculture
         Gene Function
         Animal Genetics and Genomics
      image:
         https://media.springernature.com/lw1200/springer-static/image/art%3A10.1038%2Fng.2462/MediaObjects/41588_2012_Article_BFng2462_Fig1_HTML.jpg
      isPartOf:
         name:Nature Genetics
         issn:
            1546-1718
            1061-4036
         volumeNumber:44
         type:
            Periodical
            PublicationVolume
      publisher:
         name:Nature Publishing Group US
         logo:
            url:https://www.springernature.com/app-sn/public/images/logo-springernature.png
            type:ImageObject
         type:Organization
      author:
            name:Steve Eyre
            affiliation:
                  name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre
                  address:
                     name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre, Manchester, UK
                     type:PostalAddress
                  type:Organization
                  name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre
                  address:
                     name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre, Manchester, UK
                     type:PostalAddress
                  type:Organization
            type:Person
            name:John Bowes
            affiliation:
                  name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre
                  address:
                     name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre, Manchester, UK
                     type:PostalAddress
                  type:Organization
                  name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre
                  address:
                     name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre, Manchester, UK
                     type:PostalAddress
                  type:Organization
            type:Person
            name:Dorothée Diogo
            affiliation:
                  name:Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School
                  address:
                     name:Division of Rheumatology, Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
                     type:PostalAddress
                  type:Organization
                  name:Brigham and Women's Hospital, Harvard Medical School
                  address:
                     name:Division of Genetics, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
                     type:PostalAddress
                  type:Organization
                  name:Program in Medical and Population Genetics, Broad Institute
                  address:
                     name:Program in Medical and Population Genetics, Broad Institute, Cambridge, USA
                     type:PostalAddress
                  type:Organization
            type:Person
            name:Annette Lee
            affiliation:
                  name:The Feinstein Institute for Medical Research, North Shore–Long Island Jewish Health System
                  address:
                     name:The Feinstein Institute for Medical Research, North Shore–Long Island Jewish Health System, Manhasset, USA
                     type:PostalAddress
                  type:Organization
            type:Person
            name:Anne Barton
            affiliation:
                  name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre
                  address:
                     name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre, Manchester, UK
                     type:PostalAddress
                  type:Organization
                  name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre
                  address:
                     name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre, Manchester, UK
                     type:PostalAddress
                  type:Organization
            type:Person
            name:Paul Martin
            affiliation:
                  name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre
                  address:
                     name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre, Manchester, UK
                     type:PostalAddress
                  type:Organization
                  name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre
                  address:
                     name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre, Manchester, UK
                     type:PostalAddress
                  type:Organization
            type:Person
            name:Alexandra Zhernakova
            affiliation:
                  name:Leiden University Medical Centre
                  address:
                     name:Department of Rheumatology, Leiden University Medical Centre, Leiden, The Netherlands
                     type:PostalAddress
                  type:Organization
                  name:University Medical Center Groningen and University of Groningen
                  address:
                     name:Department of Genetics, University Medical Center Groningen and University of Groningen, Groningen, The Netherlands
                     type:PostalAddress
                  type:Organization
            type:Person
            name:Eli Stahl
            affiliation:
                  name:Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School
                  address:
                     name:Division of Rheumatology, Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
                     type:PostalAddress
                  type:Organization
                  name:Brigham and Women's Hospital, Harvard Medical School
                  address:
                     name:Division of Genetics, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
                     type:PostalAddress
                  type:Organization
                  name:Program in Medical and Population Genetics, Broad Institute
                  address:
                     name:Program in Medical and Population Genetics, Broad Institute, Cambridge, USA
                     type:PostalAddress
                  type:Organization
            type:Person
            name:Sebastien Viatte
            affiliation:
                  name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre
                  address:
                     name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre, Manchester, UK
                     type:PostalAddress
                  type:Organization
                  name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre
                  address:
                     name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre, Manchester, UK
                     type:PostalAddress
                  type:Organization
            type:Person
            name:Kate McAllister
            affiliation:
                  name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre
                  address:
                     name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre, Manchester, UK
                     type:PostalAddress
                  type:Organization
                  name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre
                  address:
                     name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre, Manchester, UK
                     type:PostalAddress
                  type:Organization
            type:Person
            name:Christopher I Amos
            affiliation:
                  name:University of Texas MD Anderson Cancer Center
                  address:
                     name:University of Texas MD Anderson Cancer Center, Houston, USA
                     type:PostalAddress
                  type:Organization
            type:Person
            name:Leonid Padyukov
            affiliation:
                  name:Rheumatology Unit, Karolinska Institutet and Karolinska University Hospital Solna
                  address:
                     name:Department of Medicine, Rheumatology Unit, Karolinska Institutet and Karolinska University Hospital Solna, Stockholm, Sweden
                     type:PostalAddress
                  type:Organization
            type:Person
            name:Rene E M Toes
            affiliation:
                  name:Leiden University Medical Centre
                  address:
                     name:Department of Rheumatology, Leiden University Medical Centre, Leiden, The Netherlands
                     type:PostalAddress
                  type:Organization
            type:Person
            name:Tom W J Huizinga
            affiliation:
                  name:Leiden University Medical Centre
                  address:
                     name:Department of Rheumatology, Leiden University Medical Centre, Leiden, The Netherlands
                     type:PostalAddress
                  type:Organization
            type:Person
            name:Cisca Wijmenga
            affiliation:
                  name:University Medical Center Groningen and University of Groningen
                  address:
                     name:Department of Genetics, University Medical Center Groningen and University of Groningen, Groningen, The Netherlands
                     type:PostalAddress
                  type:Organization
            type:Person
            name:Gosia Trynka
            affiliation:
                  name:Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School
                  address:
                     name:Division of Rheumatology, Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
                     type:PostalAddress
                  type:Organization
                  name:Brigham and Women's Hospital, Harvard Medical School
                  address:
                     name:Division of Genetics, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
                     type:PostalAddress
                  type:Organization
                  name:Program in Medical and Population Genetics, Broad Institute
                  address:
                     name:Program in Medical and Population Genetics, Broad Institute, Cambridge, USA
                     type:PostalAddress
                  type:Organization
                  name:University Medical Center Groningen and University of Groningen
                  address:
                     name:Department of Genetics, University Medical Center Groningen and University of Groningen, Groningen, The Netherlands
                     type:PostalAddress
                  type:Organization
            type:Person
            name:Lude Franke
            affiliation:
                  name:University Medical Center Groningen and University of Groningen
                  address:
                     name:Department of Genetics, University Medical Center Groningen and University of Groningen, Groningen, The Netherlands
                     type:PostalAddress
                  type:Organization
            type:Person
            name:Harm-Jan Westra
            affiliation:
                  name:University Medical Center Groningen and University of Groningen
                  address:
                     name:Department of Genetics, University Medical Center Groningen and University of Groningen, Groningen, The Netherlands
                     type:PostalAddress
                  type:Organization
            type:Person
            name:Lars Alfredsson
            affiliation:
                  name:Karolinska Institutet
                  address:
                     name:Department of Environmental Medicine, Karolinska Institutet, Stockholm, Sweden
                     type:PostalAddress
                  type:Organization
            type:Person
            name:Xinli Hu
            affiliation:
                  name:Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School
                  address:
                     name:Division of Rheumatology, Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
                     type:PostalAddress
                  type:Organization
                  name:Brigham and Women's Hospital, Harvard Medical School
                  address:
                     name:Division of Genetics, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
                     type:PostalAddress
                  type:Organization
                  name:Program in Medical and Population Genetics, Broad Institute
                  address:
                     name:Program in Medical and Population Genetics, Broad Institute, Cambridge, USA
                     type:PostalAddress
                  type:Organization
                  name:Harvard–Massachusetts Institute of Technology Division of Health Sciences and Technology
                  address:
                     name:Harvard–Massachusetts Institute of Technology Division of Health Sciences and Technology, Boston, USA
                     type:PostalAddress
                  type:Organization
            type:Person
            name:Cynthia Sandor
            affiliation:
                  name:Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School
                  address:
                     name:Division of Rheumatology, Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
                     type:PostalAddress
                  type:Organization
                  name:Brigham and Women's Hospital, Harvard Medical School
                  address:
                     name:Division of Genetics, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
                     type:PostalAddress
                  type:Organization
                  name:Program in Medical and Population Genetics, Broad Institute
                  address:
                     name:Program in Medical and Population Genetics, Broad Institute, Cambridge, USA
                     type:PostalAddress
                  type:Organization
            type:Person
            name:Paul I W de Bakker
            affiliation:
                  name:Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School
                  address:
                     name:Division of Rheumatology, Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
                     type:PostalAddress
                  type:Organization
                  name:Brigham and Women's Hospital, Harvard Medical School
                  address:
                     name:Division of Genetics, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
                     type:PostalAddress
                  type:Organization
                  name:Program in Medical and Population Genetics, Broad Institute
                  address:
                     name:Program in Medical and Population Genetics, Broad Institute, Cambridge, USA
                     type:PostalAddress
                  type:Organization
                  name:University Medical Center Utrecht
                  address:
                     name:Department of Epidemiology, University Medical Center Utrecht, Utrecht, The Netherlands
                     type:PostalAddress
                  type:Organization
                  name:University Medical Center Utrecht
                  address:
                     name:Department of Medical Genetics, University Medical Center Utrecht, Utrecht, The Netherlands
                     type:PostalAddress
                  type:Organization
            type:Person
            name:Sonia Davila
            affiliation:
                  name:Genome Institute of Singapore
                  address:
                     name:Division of Human Genetics, Genome Institute of Singapore, Singapore
                     type:PostalAddress
                  type:Organization
            type:Person
            name:Chiea Chuen Khor
            affiliation:
                  name:Genome Institute of Singapore
                  address:
                     name:Division of Human Genetics, Genome Institute of Singapore, Singapore
                     type:PostalAddress
                  type:Organization
            type:Person
            name:Khai Koon Heng
            affiliation:
                  name:Genome Institute of Singapore
                  address:
                     name:Division of Human Genetics, Genome Institute of Singapore, Singapore
                     type:PostalAddress
                  type:Organization
            type:Person
            name:Robert Andrews
            affiliation:
                  name:The Wellcome Trust Sanger Institute
                  address:
                     name:The Wellcome Trust Sanger Institute, Cambridge, UK
                     type:PostalAddress
                  type:Organization
            type:Person
            name:Sarah Edkins
            affiliation:
                  name:The Wellcome Trust Sanger Institute
                  address:
                     name:The Wellcome Trust Sanger Institute, Cambridge, UK
                     type:PostalAddress
                  type:Organization
            type:Person
            name:Sarah E Hunt
            affiliation:
                  name:The Wellcome Trust Sanger Institute
                  address:
                     name:The Wellcome Trust Sanger Institute, Cambridge, UK
                     type:PostalAddress
                  type:Organization
            type:Person
            name:Cordelia Langford
            affiliation:
                  name:The Wellcome Trust Sanger Institute
                  address:
                     name:The Wellcome Trust Sanger Institute, Cambridge, UK
                     type:PostalAddress
                  type:Organization
            type:Person
            name:Deborah Symmons
            affiliation:
                  name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre
                  address:
                     name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre, Manchester, UK
                     type:PostalAddress
                  type:Organization
                  name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre
                  address:
                     name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre, Manchester, UK
                     type:PostalAddress
                  type:Organization
            type:Person
            name:Pat Concannon
            affiliation:
                  name:Center for Public Health Genomics, University of Virginia
                  address:
                     name:Center for Public Health Genomics, University of Virginia, Charlottesville, USA
                     type:PostalAddress
                  type:Organization
            type:Person
            name:Suna Onengut-Gumuscu
            affiliation:
                  name:Center for Public Health Genomics, University of Virginia
                  address:
                     name:Center for Public Health Genomics, University of Virginia, Charlottesville, USA
                     type:PostalAddress
                  type:Organization
            type:Person
            name:Stephen S Rich
            affiliation:
                  name:Center for Public Health Genomics, University of Virginia
                  address:
                     name:Center for Public Health Genomics, University of Virginia, Charlottesville, USA
                     type:PostalAddress
                  type:Organization
            type:Person
            name:Panos Deloukas
            affiliation:
                  name:The Wellcome Trust Sanger Institute
                  address:
                     name:The Wellcome Trust Sanger Institute, Cambridge, UK
                     type:PostalAddress
                  type:Organization
            type:Person
            name:Miguel A Gonzalez-Gay
            affiliation:
                  name:Hospital Marques de Valdecilla, Instituto de Fundación e Investigación Marqués de Valdecilla
                  address:
                     name:Department of Rheumatology, Hospital Marques de Valdecilla, Instituto de Fundación e Investigación Marqués de Valdecilla, Santander, Spain
                     type:PostalAddress
                  type:Organization
            type:Person
            name:Luis Rodriguez-Rodriguez
            affiliation:
                  name:Hospital Clinico San Carlos
                  address:
                     name:Hospital Clinico San Carlos, Madrid, Spain
                     type:PostalAddress
                  type:Organization
            type:Person
            name:Lisbeth Ärlsetig
            affiliation:
                  name:UmeÃ¥ University
                  address:
                     name:Department of Public Health and Clinical Medicine, UmeÃ¥ University, UmeÃ¥, Sweden
                     type:PostalAddress
                  type:Organization
                  name:UmeÃ¥ University
                  address:
                     name:Department of Rheumatology, UmeÃ¥ University, UmeÃ¥, Sweden
                     type:PostalAddress
                  type:Organization
            type:Person
            name:Javier Martin
            affiliation:
                  name:El Instituto de Parasitología y Biomedicina 'López-Neyra'–Consejo Superior de Investigaciones Científicas, Avenida del Conocimiento s/n
                  address:
                     name:El Instituto de Parasitología y Biomedicina 'López-Neyra'–Consejo Superior de Investigaciones Científicas, Avenida del Conocimiento s/n, Granada, Spain
                     type:PostalAddress
                  type:Organization
            type:Person
            name:Solbritt Rantapää-Dahlqvist
            affiliation:
                  name:UmeÃ¥ University
                  address:
                     name:Department of Public Health and Clinical Medicine, UmeÃ¥ University, UmeÃ¥, Sweden
                     type:PostalAddress
                  type:Organization
                  name:UmeÃ¥ University
                  address:
                     name:Department of Rheumatology, UmeÃ¥ University, UmeÃ¥, Sweden
                     type:PostalAddress
                  type:Organization
            type:Person
            name:Robert M Plenge
            affiliation:
                  name:Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School
                  address:
                     name:Division of Rheumatology, Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
                     type:PostalAddress
                  type:Organization
                  name:Brigham and Women's Hospital, Harvard Medical School
                  address:
                     name:Division of Genetics, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
                     type:PostalAddress
                  type:Organization
                  name:Program in Medical and Population Genetics, Broad Institute
                  address:
                     name:Program in Medical and Population Genetics, Broad Institute, Cambridge, USA
                     type:PostalAddress
                  type:Organization
            type:Person
            name:Soumya Raychaudhuri
            affiliation:
                  name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre
                  address:
                     name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre, Manchester, UK
                     type:PostalAddress
                  type:Organization
                  name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre
                  address:
                     name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre, Manchester, UK
                     type:PostalAddress
                  type:Organization
                  name:Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School
                  address:
                     name:Division of Rheumatology, Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
                     type:PostalAddress
                  type:Organization
                  name:Brigham and Women's Hospital, Harvard Medical School
                  address:
                     name:Division of Genetics, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
                     type:PostalAddress
                  type:Organization
                  name:Program in Medical and Population Genetics, Broad Institute
                  address:
                     name:Program in Medical and Population Genetics, Broad Institute, Cambridge, USA
                     type:PostalAddress
                  type:Organization
            type:Person
            name:Lars Klareskog
            affiliation:
                  name:Rheumatology Unit, Karolinska Institutet and Karolinska University Hospital Solna
                  address:
                     name:Department of Medicine, Rheumatology Unit, Karolinska Institutet and Karolinska University Hospital Solna, Stockholm, Sweden
                     type:PostalAddress
                  type:Organization
            type:Person
            name:Peter K Gregersen
            affiliation:
                  name:The Feinstein Institute for Medical Research, North Shore–Long Island Jewish Health System
                  address:
                     name:The Feinstein Institute for Medical Research, North Shore–Long Island Jewish Health System, Manhasset, USA
                     type:PostalAddress
                  type:Organization
            type:Person
            name:Jane Worthington
            affiliation:
                  name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre
                  address:
                     name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre, Manchester, UK
                     type:PostalAddress
                  type:Organization
                  name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre
                  address:
                     name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre, Manchester, UK
                     type:PostalAddress
                  type:Organization
            email:[email protected]
            type:Person
      isAccessibleForFree:
      hasPart:
         isAccessibleForFree:
         cssSelector:.main-content
         type:WebPageElement
["Periodical","PublicationVolume"]:
      name:Nature Genetics
      issn:
         1546-1718
         1061-4036
      volumeNumber:44
Organization:
      name:Nature Publishing Group US
      logo:
         url:https://www.springernature.com/app-sn/public/images/logo-springernature.png
         type:ImageObject
      name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre
      address:
         name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre, Manchester, UK
         type:PostalAddress
      name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre
      address:
         name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre, Manchester, UK
         type:PostalAddress
      name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre
      address:
         name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre, Manchester, UK
         type:PostalAddress
      name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre
      address:
         name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre, Manchester, UK
         type:PostalAddress
      name:Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School
      address:
         name:Division of Rheumatology, Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
         type:PostalAddress
      name:Brigham and Women's Hospital, Harvard Medical School
      address:
         name:Division of Genetics, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
         type:PostalAddress
      name:Program in Medical and Population Genetics, Broad Institute
      address:
         name:Program in Medical and Population Genetics, Broad Institute, Cambridge, USA
         type:PostalAddress
      name:The Feinstein Institute for Medical Research, North Shore–Long Island Jewish Health System
      address:
         name:The Feinstein Institute for Medical Research, North Shore–Long Island Jewish Health System, Manhasset, USA
         type:PostalAddress
      name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre
      address:
         name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre, Manchester, UK
         type:PostalAddress
      name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre
      address:
         name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre, Manchester, UK
         type:PostalAddress
      name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre
      address:
         name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre, Manchester, UK
         type:PostalAddress
      name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre
      address:
         name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre, Manchester, UK
         type:PostalAddress
      name:Leiden University Medical Centre
      address:
         name:Department of Rheumatology, Leiden University Medical Centre, Leiden, The Netherlands
         type:PostalAddress
      name:University Medical Center Groningen and University of Groningen
      address:
         name:Department of Genetics, University Medical Center Groningen and University of Groningen, Groningen, The Netherlands
         type:PostalAddress
      name:Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School
      address:
         name:Division of Rheumatology, Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
         type:PostalAddress
      name:Brigham and Women's Hospital, Harvard Medical School
      address:
         name:Division of Genetics, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
         type:PostalAddress
      name:Program in Medical and Population Genetics, Broad Institute
      address:
         name:Program in Medical and Population Genetics, Broad Institute, Cambridge, USA
         type:PostalAddress
      name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre
      address:
         name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre, Manchester, UK
         type:PostalAddress
      name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre
      address:
         name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre, Manchester, UK
         type:PostalAddress
      name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre
      address:
         name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre, Manchester, UK
         type:PostalAddress
      name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre
      address:
         name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre, Manchester, UK
         type:PostalAddress
      name:University of Texas MD Anderson Cancer Center
      address:
         name:University of Texas MD Anderson Cancer Center, Houston, USA
         type:PostalAddress
      name:Rheumatology Unit, Karolinska Institutet and Karolinska University Hospital Solna
      address:
         name:Department of Medicine, Rheumatology Unit, Karolinska Institutet and Karolinska University Hospital Solna, Stockholm, Sweden
         type:PostalAddress
      name:Leiden University Medical Centre
      address:
         name:Department of Rheumatology, Leiden University Medical Centre, Leiden, The Netherlands
         type:PostalAddress
      name:Leiden University Medical Centre
      address:
         name:Department of Rheumatology, Leiden University Medical Centre, Leiden, The Netherlands
         type:PostalAddress
      name:University Medical Center Groningen and University of Groningen
      address:
         name:Department of Genetics, University Medical Center Groningen and University of Groningen, Groningen, The Netherlands
         type:PostalAddress
      name:Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School
      address:
         name:Division of Rheumatology, Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
         type:PostalAddress
      name:Brigham and Women's Hospital, Harvard Medical School
      address:
         name:Division of Genetics, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
         type:PostalAddress
      name:Program in Medical and Population Genetics, Broad Institute
      address:
         name:Program in Medical and Population Genetics, Broad Institute, Cambridge, USA
         type:PostalAddress
      name:University Medical Center Groningen and University of Groningen
      address:
         name:Department of Genetics, University Medical Center Groningen and University of Groningen, Groningen, The Netherlands
         type:PostalAddress
      name:University Medical Center Groningen and University of Groningen
      address:
         name:Department of Genetics, University Medical Center Groningen and University of Groningen, Groningen, The Netherlands
         type:PostalAddress
      name:University Medical Center Groningen and University of Groningen
      address:
         name:Department of Genetics, University Medical Center Groningen and University of Groningen, Groningen, The Netherlands
         type:PostalAddress
      name:Karolinska Institutet
      address:
         name:Department of Environmental Medicine, Karolinska Institutet, Stockholm, Sweden
         type:PostalAddress
      name:Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School
      address:
         name:Division of Rheumatology, Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
         type:PostalAddress
      name:Brigham and Women's Hospital, Harvard Medical School
      address:
         name:Division of Genetics, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
         type:PostalAddress
      name:Program in Medical and Population Genetics, Broad Institute
      address:
         name:Program in Medical and Population Genetics, Broad Institute, Cambridge, USA
         type:PostalAddress
      name:Harvard–Massachusetts Institute of Technology Division of Health Sciences and Technology
      address:
         name:Harvard–Massachusetts Institute of Technology Division of Health Sciences and Technology, Boston, USA
         type:PostalAddress
      name:Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School
      address:
         name:Division of Rheumatology, Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
         type:PostalAddress
      name:Brigham and Women's Hospital, Harvard Medical School
      address:
         name:Division of Genetics, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
         type:PostalAddress
      name:Program in Medical and Population Genetics, Broad Institute
      address:
         name:Program in Medical and Population Genetics, Broad Institute, Cambridge, USA
         type:PostalAddress
      name:Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School
      address:
         name:Division of Rheumatology, Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
         type:PostalAddress
      name:Brigham and Women's Hospital, Harvard Medical School
      address:
         name:Division of Genetics, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
         type:PostalAddress
      name:Program in Medical and Population Genetics, Broad Institute
      address:
         name:Program in Medical and Population Genetics, Broad Institute, Cambridge, USA
         type:PostalAddress
      name:University Medical Center Utrecht
      address:
         name:Department of Epidemiology, University Medical Center Utrecht, Utrecht, The Netherlands
         type:PostalAddress
      name:University Medical Center Utrecht
      address:
         name:Department of Medical Genetics, University Medical Center Utrecht, Utrecht, The Netherlands
         type:PostalAddress
      name:Genome Institute of Singapore
      address:
         name:Division of Human Genetics, Genome Institute of Singapore, Singapore
         type:PostalAddress
      name:Genome Institute of Singapore
      address:
         name:Division of Human Genetics, Genome Institute of Singapore, Singapore
         type:PostalAddress
      name:Genome Institute of Singapore
      address:
         name:Division of Human Genetics, Genome Institute of Singapore, Singapore
         type:PostalAddress
      name:The Wellcome Trust Sanger Institute
      address:
         name:The Wellcome Trust Sanger Institute, Cambridge, UK
         type:PostalAddress
      name:The Wellcome Trust Sanger Institute
      address:
         name:The Wellcome Trust Sanger Institute, Cambridge, UK
         type:PostalAddress
      name:The Wellcome Trust Sanger Institute
      address:
         name:The Wellcome Trust Sanger Institute, Cambridge, UK
         type:PostalAddress
      name:The Wellcome Trust Sanger Institute
      address:
         name:The Wellcome Trust Sanger Institute, Cambridge, UK
         type:PostalAddress
      name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre
      address:
         name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre, Manchester, UK
         type:PostalAddress
      name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre
      address:
         name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre, Manchester, UK
         type:PostalAddress
      name:Center for Public Health Genomics, University of Virginia
      address:
         name:Center for Public Health Genomics, University of Virginia, Charlottesville, USA
         type:PostalAddress
      name:Center for Public Health Genomics, University of Virginia
      address:
         name:Center for Public Health Genomics, University of Virginia, Charlottesville, USA
         type:PostalAddress
      name:Center for Public Health Genomics, University of Virginia
      address:
         name:Center for Public Health Genomics, University of Virginia, Charlottesville, USA
         type:PostalAddress
      name:The Wellcome Trust Sanger Institute
      address:
         name:The Wellcome Trust Sanger Institute, Cambridge, UK
         type:PostalAddress
      name:Hospital Marques de Valdecilla, Instituto de Fundación e Investigación Marqués de Valdecilla
      address:
         name:Department of Rheumatology, Hospital Marques de Valdecilla, Instituto de Fundación e Investigación Marqués de Valdecilla, Santander, Spain
         type:PostalAddress
      name:Hospital Clinico San Carlos
      address:
         name:Hospital Clinico San Carlos, Madrid, Spain
         type:PostalAddress
      name:UmeÃ¥ University
      address:
         name:Department of Public Health and Clinical Medicine, UmeÃ¥ University, UmeÃ¥, Sweden
         type:PostalAddress
      name:UmeÃ¥ University
      address:
         name:Department of Rheumatology, UmeÃ¥ University, UmeÃ¥, Sweden
         type:PostalAddress
      name:El Instituto de Parasitología y Biomedicina 'López-Neyra'–Consejo Superior de Investigaciones Científicas, Avenida del Conocimiento s/n
      address:
         name:El Instituto de Parasitología y Biomedicina 'López-Neyra'–Consejo Superior de Investigaciones Científicas, Avenida del Conocimiento s/n, Granada, Spain
         type:PostalAddress
      name:UmeÃ¥ University
      address:
         name:Department of Public Health and Clinical Medicine, UmeÃ¥ University, UmeÃ¥, Sweden
         type:PostalAddress
      name:UmeÃ¥ University
      address:
         name:Department of Rheumatology, UmeÃ¥ University, UmeÃ¥, Sweden
         type:PostalAddress
      name:Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School
      address:
         name:Division of Rheumatology, Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
         type:PostalAddress
      name:Brigham and Women's Hospital, Harvard Medical School
      address:
         name:Division of Genetics, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
         type:PostalAddress
      name:Program in Medical and Population Genetics, Broad Institute
      address:
         name:Program in Medical and Population Genetics, Broad Institute, Cambridge, USA
         type:PostalAddress
      name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre
      address:
         name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre, Manchester, UK
         type:PostalAddress
      name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre
      address:
         name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre, Manchester, UK
         type:PostalAddress
      name:Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School
      address:
         name:Division of Rheumatology, Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
         type:PostalAddress
      name:Brigham and Women's Hospital, Harvard Medical School
      address:
         name:Division of Genetics, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
         type:PostalAddress
      name:Program in Medical and Population Genetics, Broad Institute
      address:
         name:Program in Medical and Population Genetics, Broad Institute, Cambridge, USA
         type:PostalAddress
      name:Rheumatology Unit, Karolinska Institutet and Karolinska University Hospital Solna
      address:
         name:Department of Medicine, Rheumatology Unit, Karolinska Institutet and Karolinska University Hospital Solna, Stockholm, Sweden
         type:PostalAddress
      name:The Feinstein Institute for Medical Research, North Shore–Long Island Jewish Health System
      address:
         name:The Feinstein Institute for Medical Research, North Shore–Long Island Jewish Health System, Manhasset, USA
         type:PostalAddress
      name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre
      address:
         name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre, Manchester, UK
         type:PostalAddress
      name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre
      address:
         name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre, Manchester, UK
         type:PostalAddress
ImageObject:
      url:https://www.springernature.com/app-sn/public/images/logo-springernature.png
Person:
      name:Steve Eyre
      affiliation:
            name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre
            address:
               name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre, Manchester, UK
               type:PostalAddress
            type:Organization
            name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre
            address:
               name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre, Manchester, UK
               type:PostalAddress
            type:Organization
      name:John Bowes
      affiliation:
            name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre
            address:
               name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre, Manchester, UK
               type:PostalAddress
            type:Organization
            name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre
            address:
               name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre, Manchester, UK
               type:PostalAddress
            type:Organization
      name:Dorothée Diogo
      affiliation:
            name:Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School
            address:
               name:Division of Rheumatology, Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
               type:PostalAddress
            type:Organization
            name:Brigham and Women's Hospital, Harvard Medical School
            address:
               name:Division of Genetics, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
               type:PostalAddress
            type:Organization
            name:Program in Medical and Population Genetics, Broad Institute
            address:
               name:Program in Medical and Population Genetics, Broad Institute, Cambridge, USA
               type:PostalAddress
            type:Organization
      name:Annette Lee
      affiliation:
            name:The Feinstein Institute for Medical Research, North Shore–Long Island Jewish Health System
            address:
               name:The Feinstein Institute for Medical Research, North Shore–Long Island Jewish Health System, Manhasset, USA
               type:PostalAddress
            type:Organization
      name:Anne Barton
      affiliation:
            name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre
            address:
               name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre, Manchester, UK
               type:PostalAddress
            type:Organization
            name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre
            address:
               name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre, Manchester, UK
               type:PostalAddress
            type:Organization
      name:Paul Martin
      affiliation:
            name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre
            address:
               name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre, Manchester, UK
               type:PostalAddress
            type:Organization
            name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre
            address:
               name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre, Manchester, UK
               type:PostalAddress
            type:Organization
      name:Alexandra Zhernakova
      affiliation:
            name:Leiden University Medical Centre
            address:
               name:Department of Rheumatology, Leiden University Medical Centre, Leiden, The Netherlands
               type:PostalAddress
            type:Organization
            name:University Medical Center Groningen and University of Groningen
            address:
               name:Department of Genetics, University Medical Center Groningen and University of Groningen, Groningen, The Netherlands
               type:PostalAddress
            type:Organization
      name:Eli Stahl
      affiliation:
            name:Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School
            address:
               name:Division of Rheumatology, Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
               type:PostalAddress
            type:Organization
            name:Brigham and Women's Hospital, Harvard Medical School
            address:
               name:Division of Genetics, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
               type:PostalAddress
            type:Organization
            name:Program in Medical and Population Genetics, Broad Institute
            address:
               name:Program in Medical and Population Genetics, Broad Institute, Cambridge, USA
               type:PostalAddress
            type:Organization
      name:Sebastien Viatte
      affiliation:
            name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre
            address:
               name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre, Manchester, UK
               type:PostalAddress
            type:Organization
            name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre
            address:
               name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre, Manchester, UK
               type:PostalAddress
            type:Organization
      name:Kate McAllister
      affiliation:
            name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre
            address:
               name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre, Manchester, UK
               type:PostalAddress
            type:Organization
            name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre
            address:
               name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre, Manchester, UK
               type:PostalAddress
            type:Organization
      name:Christopher I Amos
      affiliation:
            name:University of Texas MD Anderson Cancer Center
            address:
               name:University of Texas MD Anderson Cancer Center, Houston, USA
               type:PostalAddress
            type:Organization
      name:Leonid Padyukov
      affiliation:
            name:Rheumatology Unit, Karolinska Institutet and Karolinska University Hospital Solna
            address:
               name:Department of Medicine, Rheumatology Unit, Karolinska Institutet and Karolinska University Hospital Solna, Stockholm, Sweden
               type:PostalAddress
            type:Organization
      name:Rene E M Toes
      affiliation:
            name:Leiden University Medical Centre
            address:
               name:Department of Rheumatology, Leiden University Medical Centre, Leiden, The Netherlands
               type:PostalAddress
            type:Organization
      name:Tom W J Huizinga
      affiliation:
            name:Leiden University Medical Centre
            address:
               name:Department of Rheumatology, Leiden University Medical Centre, Leiden, The Netherlands
               type:PostalAddress
            type:Organization
      name:Cisca Wijmenga
      affiliation:
            name:University Medical Center Groningen and University of Groningen
            address:
               name:Department of Genetics, University Medical Center Groningen and University of Groningen, Groningen, The Netherlands
               type:PostalAddress
            type:Organization
      name:Gosia Trynka
      affiliation:
            name:Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School
            address:
               name:Division of Rheumatology, Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
               type:PostalAddress
            type:Organization
            name:Brigham and Women's Hospital, Harvard Medical School
            address:
               name:Division of Genetics, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
               type:PostalAddress
            type:Organization
            name:Program in Medical and Population Genetics, Broad Institute
            address:
               name:Program in Medical and Population Genetics, Broad Institute, Cambridge, USA
               type:PostalAddress
            type:Organization
            name:University Medical Center Groningen and University of Groningen
            address:
               name:Department of Genetics, University Medical Center Groningen and University of Groningen, Groningen, The Netherlands
               type:PostalAddress
            type:Organization
      name:Lude Franke
      affiliation:
            name:University Medical Center Groningen and University of Groningen
            address:
               name:Department of Genetics, University Medical Center Groningen and University of Groningen, Groningen, The Netherlands
               type:PostalAddress
            type:Organization
      name:Harm-Jan Westra
      affiliation:
            name:University Medical Center Groningen and University of Groningen
            address:
               name:Department of Genetics, University Medical Center Groningen and University of Groningen, Groningen, The Netherlands
               type:PostalAddress
            type:Organization
      name:Lars Alfredsson
      affiliation:
            name:Karolinska Institutet
            address:
               name:Department of Environmental Medicine, Karolinska Institutet, Stockholm, Sweden
               type:PostalAddress
            type:Organization
      name:Xinli Hu
      affiliation:
            name:Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School
            address:
               name:Division of Rheumatology, Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
               type:PostalAddress
            type:Organization
            name:Brigham and Women's Hospital, Harvard Medical School
            address:
               name:Division of Genetics, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
               type:PostalAddress
            type:Organization
            name:Program in Medical and Population Genetics, Broad Institute
            address:
               name:Program in Medical and Population Genetics, Broad Institute, Cambridge, USA
               type:PostalAddress
            type:Organization
            name:Harvard–Massachusetts Institute of Technology Division of Health Sciences and Technology
            address:
               name:Harvard–Massachusetts Institute of Technology Division of Health Sciences and Technology, Boston, USA
               type:PostalAddress
            type:Organization
      name:Cynthia Sandor
      affiliation:
            name:Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School
            address:
               name:Division of Rheumatology, Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
               type:PostalAddress
            type:Organization
            name:Brigham and Women's Hospital, Harvard Medical School
            address:
               name:Division of Genetics, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
               type:PostalAddress
            type:Organization
            name:Program in Medical and Population Genetics, Broad Institute
            address:
               name:Program in Medical and Population Genetics, Broad Institute, Cambridge, USA
               type:PostalAddress
            type:Organization
      name:Paul I W de Bakker
      affiliation:
            name:Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School
            address:
               name:Division of Rheumatology, Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
               type:PostalAddress
            type:Organization
            name:Brigham and Women's Hospital, Harvard Medical School
            address:
               name:Division of Genetics, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
               type:PostalAddress
            type:Organization
            name:Program in Medical and Population Genetics, Broad Institute
            address:
               name:Program in Medical and Population Genetics, Broad Institute, Cambridge, USA
               type:PostalAddress
            type:Organization
            name:University Medical Center Utrecht
            address:
               name:Department of Epidemiology, University Medical Center Utrecht, Utrecht, The Netherlands
               type:PostalAddress
            type:Organization
            name:University Medical Center Utrecht
            address:
               name:Department of Medical Genetics, University Medical Center Utrecht, Utrecht, The Netherlands
               type:PostalAddress
            type:Organization
      name:Sonia Davila
      affiliation:
            name:Genome Institute of Singapore
            address:
               name:Division of Human Genetics, Genome Institute of Singapore, Singapore
               type:PostalAddress
            type:Organization
      name:Chiea Chuen Khor
      affiliation:
            name:Genome Institute of Singapore
            address:
               name:Division of Human Genetics, Genome Institute of Singapore, Singapore
               type:PostalAddress
            type:Organization
      name:Khai Koon Heng
      affiliation:
            name:Genome Institute of Singapore
            address:
               name:Division of Human Genetics, Genome Institute of Singapore, Singapore
               type:PostalAddress
            type:Organization
      name:Robert Andrews
      affiliation:
            name:The Wellcome Trust Sanger Institute
            address:
               name:The Wellcome Trust Sanger Institute, Cambridge, UK
               type:PostalAddress
            type:Organization
      name:Sarah Edkins
      affiliation:
            name:The Wellcome Trust Sanger Institute
            address:
               name:The Wellcome Trust Sanger Institute, Cambridge, UK
               type:PostalAddress
            type:Organization
      name:Sarah E Hunt
      affiliation:
            name:The Wellcome Trust Sanger Institute
            address:
               name:The Wellcome Trust Sanger Institute, Cambridge, UK
               type:PostalAddress
            type:Organization
      name:Cordelia Langford
      affiliation:
            name:The Wellcome Trust Sanger Institute
            address:
               name:The Wellcome Trust Sanger Institute, Cambridge, UK
               type:PostalAddress
            type:Organization
      name:Deborah Symmons
      affiliation:
            name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre
            address:
               name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre, Manchester, UK
               type:PostalAddress
            type:Organization
            name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre
            address:
               name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre, Manchester, UK
               type:PostalAddress
            type:Organization
      name:Pat Concannon
      affiliation:
            name:Center for Public Health Genomics, University of Virginia
            address:
               name:Center for Public Health Genomics, University of Virginia, Charlottesville, USA
               type:PostalAddress
            type:Organization
      name:Suna Onengut-Gumuscu
      affiliation:
            name:Center for Public Health Genomics, University of Virginia
            address:
               name:Center for Public Health Genomics, University of Virginia, Charlottesville, USA
               type:PostalAddress
            type:Organization
      name:Stephen S Rich
      affiliation:
            name:Center for Public Health Genomics, University of Virginia
            address:
               name:Center for Public Health Genomics, University of Virginia, Charlottesville, USA
               type:PostalAddress
            type:Organization
      name:Panos Deloukas
      affiliation:
            name:The Wellcome Trust Sanger Institute
            address:
               name:The Wellcome Trust Sanger Institute, Cambridge, UK
               type:PostalAddress
            type:Organization
      name:Miguel A Gonzalez-Gay
      affiliation:
            name:Hospital Marques de Valdecilla, Instituto de Fundación e Investigación Marqués de Valdecilla
            address:
               name:Department of Rheumatology, Hospital Marques de Valdecilla, Instituto de Fundación e Investigación Marqués de Valdecilla, Santander, Spain
               type:PostalAddress
            type:Organization
      name:Luis Rodriguez-Rodriguez
      affiliation:
            name:Hospital Clinico San Carlos
            address:
               name:Hospital Clinico San Carlos, Madrid, Spain
               type:PostalAddress
            type:Organization
      name:Lisbeth Ärlsetig
      affiliation:
            name:UmeÃ¥ University
            address:
               name:Department of Public Health and Clinical Medicine, UmeÃ¥ University, UmeÃ¥, Sweden
               type:PostalAddress
            type:Organization
            name:UmeÃ¥ University
            address:
               name:Department of Rheumatology, UmeÃ¥ University, UmeÃ¥, Sweden
               type:PostalAddress
            type:Organization
      name:Javier Martin
      affiliation:
            name:El Instituto de Parasitología y Biomedicina 'López-Neyra'–Consejo Superior de Investigaciones Científicas, Avenida del Conocimiento s/n
            address:
               name:El Instituto de Parasitología y Biomedicina 'López-Neyra'–Consejo Superior de Investigaciones Científicas, Avenida del Conocimiento s/n, Granada, Spain
               type:PostalAddress
            type:Organization
      name:Solbritt Rantapää-Dahlqvist
      affiliation:
            name:UmeÃ¥ University
            address:
               name:Department of Public Health and Clinical Medicine, UmeÃ¥ University, UmeÃ¥, Sweden
               type:PostalAddress
            type:Organization
            name:UmeÃ¥ University
            address:
               name:Department of Rheumatology, UmeÃ¥ University, UmeÃ¥, Sweden
               type:PostalAddress
            type:Organization
      name:Robert M Plenge
      affiliation:
            name:Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School
            address:
               name:Division of Rheumatology, Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
               type:PostalAddress
            type:Organization
            name:Brigham and Women's Hospital, Harvard Medical School
            address:
               name:Division of Genetics, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
               type:PostalAddress
            type:Organization
            name:Program in Medical and Population Genetics, Broad Institute
            address:
               name:Program in Medical and Population Genetics, Broad Institute, Cambridge, USA
               type:PostalAddress
            type:Organization
      name:Soumya Raychaudhuri
      affiliation:
            name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre
            address:
               name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre, Manchester, UK
               type:PostalAddress
            type:Organization
            name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre
            address:
               name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre, Manchester, UK
               type:PostalAddress
            type:Organization
            name:Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School
            address:
               name:Division of Rheumatology, Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
               type:PostalAddress
            type:Organization
            name:Brigham and Women's Hospital, Harvard Medical School
            address:
               name:Division of Genetics, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
               type:PostalAddress
            type:Organization
            name:Program in Medical and Population Genetics, Broad Institute
            address:
               name:Program in Medical and Population Genetics, Broad Institute, Cambridge, USA
               type:PostalAddress
            type:Organization
      name:Lars Klareskog
      affiliation:
            name:Rheumatology Unit, Karolinska Institutet and Karolinska University Hospital Solna
            address:
               name:Department of Medicine, Rheumatology Unit, Karolinska Institutet and Karolinska University Hospital Solna, Stockholm, Sweden
               type:PostalAddress
            type:Organization
      name:Peter K Gregersen
      affiliation:
            name:The Feinstein Institute for Medical Research, North Shore–Long Island Jewish Health System
            address:
               name:The Feinstein Institute for Medical Research, North Shore–Long Island Jewish Health System, Manhasset, USA
               type:PostalAddress
            type:Organization
      name:Jane Worthington
      affiliation:
            name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre
            address:
               name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre, Manchester, UK
               type:PostalAddress
            type:Organization
            name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre
            address:
               name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre, Manchester, UK
               type:PostalAddress
            type:Organization
      email:[email protected]
PostalAddress:
      name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre, Manchester, UK
      name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre, Manchester, UK
      name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre, Manchester, UK
      name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre, Manchester, UK
      name:Division of Rheumatology, Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
      name:Division of Genetics, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
      name:Program in Medical and Population Genetics, Broad Institute, Cambridge, USA
      name:The Feinstein Institute for Medical Research, North Shore–Long Island Jewish Health System, Manhasset, USA
      name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre, Manchester, UK
      name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre, Manchester, UK
      name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre, Manchester, UK
      name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre, Manchester, UK
      name:Department of Rheumatology, Leiden University Medical Centre, Leiden, The Netherlands
      name:Department of Genetics, University Medical Center Groningen and University of Groningen, Groningen, The Netherlands
      name:Division of Rheumatology, Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
      name:Division of Genetics, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
      name:Program in Medical and Population Genetics, Broad Institute, Cambridge, USA
      name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre, Manchester, UK
      name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre, Manchester, UK
      name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre, Manchester, UK
      name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre, Manchester, UK
      name:University of Texas MD Anderson Cancer Center, Houston, USA
      name:Department of Medicine, Rheumatology Unit, Karolinska Institutet and Karolinska University Hospital Solna, Stockholm, Sweden
      name:Department of Rheumatology, Leiden University Medical Centre, Leiden, The Netherlands
      name:Department of Rheumatology, Leiden University Medical Centre, Leiden, The Netherlands
      name:Department of Genetics, University Medical Center Groningen and University of Groningen, Groningen, The Netherlands
      name:Division of Rheumatology, Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
      name:Division of Genetics, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
      name:Program in Medical and Population Genetics, Broad Institute, Cambridge, USA
      name:Department of Genetics, University Medical Center Groningen and University of Groningen, Groningen, The Netherlands
      name:Department of Genetics, University Medical Center Groningen and University of Groningen, Groningen, The Netherlands
      name:Department of Genetics, University Medical Center Groningen and University of Groningen, Groningen, The Netherlands
      name:Department of Environmental Medicine, Karolinska Institutet, Stockholm, Sweden
      name:Division of Rheumatology, Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
      name:Division of Genetics, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
      name:Program in Medical and Population Genetics, Broad Institute, Cambridge, USA
      name:Harvard–Massachusetts Institute of Technology Division of Health Sciences and Technology, Boston, USA
      name:Division of Rheumatology, Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
      name:Division of Genetics, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
      name:Program in Medical and Population Genetics, Broad Institute, Cambridge, USA
      name:Division of Rheumatology, Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
      name:Division of Genetics, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
      name:Program in Medical and Population Genetics, Broad Institute, Cambridge, USA
      name:Department of Epidemiology, University Medical Center Utrecht, Utrecht, The Netherlands
      name:Department of Medical Genetics, University Medical Center Utrecht, Utrecht, The Netherlands
      name:Division of Human Genetics, Genome Institute of Singapore, Singapore
      name:Division of Human Genetics, Genome Institute of Singapore, Singapore
      name:Division of Human Genetics, Genome Institute of Singapore, Singapore
      name:The Wellcome Trust Sanger Institute, Cambridge, UK
      name:The Wellcome Trust Sanger Institute, Cambridge, UK
      name:The Wellcome Trust Sanger Institute, Cambridge, UK
      name:The Wellcome Trust Sanger Institute, Cambridge, UK
      name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre, Manchester, UK
      name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre, Manchester, UK
      name:Center for Public Health Genomics, University of Virginia, Charlottesville, USA
      name:Center for Public Health Genomics, University of Virginia, Charlottesville, USA
      name:Center for Public Health Genomics, University of Virginia, Charlottesville, USA
      name:The Wellcome Trust Sanger Institute, Cambridge, UK
      name:Department of Rheumatology, Hospital Marques de Valdecilla, Instituto de Fundación e Investigación Marqués de Valdecilla, Santander, Spain
      name:Hospital Clinico San Carlos, Madrid, Spain
      name:Department of Public Health and Clinical Medicine, UmeÃ¥ University, UmeÃ¥, Sweden
      name:Department of Rheumatology, UmeÃ¥ University, UmeÃ¥, Sweden
      name:El Instituto de Parasitología y Biomedicina 'López-Neyra'–Consejo Superior de Investigaciones Científicas, Avenida del Conocimiento s/n, Granada, Spain
      name:Department of Public Health and Clinical Medicine, UmeÃ¥ University, UmeÃ¥, Sweden
      name:Department of Rheumatology, UmeÃ¥ University, UmeÃ¥, Sweden
      name:Division of Rheumatology, Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
      name:Division of Genetics, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
      name:Program in Medical and Population Genetics, Broad Institute, Cambridge, USA
      name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre, Manchester, UK
      name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre, Manchester, UK
      name:Division of Rheumatology, Immunology and Allergy, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
      name:Division of Genetics, Brigham and Women's Hospital, Harvard Medical School, Boston, USA
      name:Program in Medical and Population Genetics, Broad Institute, Cambridge, USA
      name:Department of Medicine, Rheumatology Unit, Karolinska Institutet and Karolinska University Hospital Solna, Stockholm, Sweden
      name:The Feinstein Institute for Medical Research, North Shore–Long Island Jewish Health System, Manhasset, USA
      name:Arthritis Research UK Epidemiology Unit, Centre for Musculoskeletal Research, University of Manchester, Manchester Academic Health Science Centre, Manchester, UK
      name:National Institute for Health Research, Manchester Musculoskeletal Biomedical Research Unit, Central Manchester University Hospitals National Health Service Foundation Trust, Manchester Academic Health Sciences Centre, Manchester, UK
WebPageElement:
      isAccessibleForFree:
      cssSelector:.main-content

Social Networks {👍}(2)

• https://www.facebook.com/NatureGenet
• https://twitter.com/NatureGenet

External Links {🔗}(147)

• How much money does https://link.springer.com/article/10.1038/ng.2462?utm_source=nature&utm_medium=referral&utm_campaign=buyArticle generate?
• Financial intake of https://doi.org/10.1038%2Fng.582
• Get to know what's the income of http://scholar.google.com/scholar_lookup?&title=Genome-wide%20association%20study%20meta-analysis%20identifies%20seven%20new%20rheumatoid%20arthritis%20risk%20loci&journal=Nat.%20Genet.&doi=10.1038%2Fng.582&volume=42&pages=508-514&publication_year=2010&author=Stahl%2CEA
• How much does https://doi.org/10.1371%2Fjournal.pgen.1002004 net monthly?
• Profit of http://scholar.google.com/scholar_lookup?&title=Meta-analysis%20of%20genome-wide%20association%20studies%20in%20celiac%20disease%20and%20rheumatoid%20arthritis%20identifies%20fourteen%20non-HLA%20shared%20loci&journal=PLoS%20Genet.&doi=10.1371%2Fjournal.pgen.1002004&volume=7&publication_year=2011&author=Zhernakova%2CA
• Get to know what's the income of https://doi.org/10.1038%2Fng.998
• Discover the revenue of http://scholar.google.com/scholar_lookup?&title=Dense%20genotyping%20identifies%20and%20localizes%20multiple%20common%20and%20rare%20variant%20association%20signals%20in%20celiac%20disease&journal=Nat.%20Genet.&doi=10.1038%2Fng.998&volume=43&pages=1193-1201&publication_year=2011&author=Trynka%2CG
• What are the earnings of https://doi.org/10.1038%2Fng.2232?
• How much profit does http://scholar.google.com/scholar_lookup?&title=Bayesian%20inference%20analyses%20of%20the%20polygenic%20architecture%20of%20rheumatoid%20arthritis&journal=Nat.%20Genet.&doi=10.1038%2Fng.2232&volume=44&pages=483-489&publication_year=2012&author=Stahl%2CEA generate?
• What's the income of https://doi.org/10.1038%2Fng.1076?
• What's the income of http://scholar.google.com/scholar_lookup?&title=Five%20amino%20acids%20in%20three%20HLA%20proteins%20explain%20most%20of%20the%20association%20between%20MHC%20and%20seropositive%20rheumatoid%20arthritis&journal=Nat.%20Genet.&doi=10.1038%2Fng.1076&volume=44&pages=291-296&publication_year=2012&author=Raychaudhuri%2CS?
• Earnings of https://doi.org/10.1136%2Fard.2009.126821
• Find out how much http://scholar.google.com/scholar_lookup?&title=A%20genome-wide%20association%20study%20suggests%20contrasting%20associations%20in%20ACPA-positive%20versus%20ACPA-negative%20rheumatoid%20arthritis&journal=Ann.%20Rheum.%20Dis.&doi=10.1136%2Fard.2009.126821&volume=70&pages=259-265&publication_year=2011&author=Padyukov%2CL earns monthly
• How much revenue does https://doi.org/10.1073%2Fpnas.0901181106 generate?
• Revenue of http://scholar.google.com/scholar_lookup?&title=Identification%20of%20IRAK1%20as%20a%20risk%20gene%20with%20critical%20role%20in%20the%20pathogenesis%20of%20systemic%20lupus%20erythematosus&journal=Proc.%20Natl.%20Acad.%20Sci.%20USA&doi=10.1073%2Fpnas.0901181106&volume=106&pages=6256-6261&publication_year=2009&author=Jacob%2CCO
• Learn about the earnings of https://doi.org/10.1038%2Fnature03479
• What's the income of http://scholar.google.com/scholar_lookup?&title=X-inactivation%20profile%20reveals%20extensive%20variability%20in%20X-linked%20gene%20expression%20in%20females&journal=Nature&doi=10.1038%2Fnature03479&volume=434&pages=400-404&publication_year=2005&author=Carrel%2CL&author=Willard%2CHF?
• How profitable is https://doi.org/10.1038%2Fng1206?
• What's the income of http://scholar.google.com/scholar_lookup?&title=Functional%20haplotypes%20of%20PADI4%2C%20encoding%20citrullinating%20enzyme%20peptidylarginine%20deiminase%204%2C%20are%20associated%20with%20rheumatoid%20arthritis&journal=Nat.%20Genet.&doi=10.1038%2Fng1206&volume=34&pages=395-402&publication_year=2003&author=Suzuki%2CA?
• How much revenue does https://doi.org/10.1016%2Fj.ajhg.2012.01.010 produce monthly?
• What's the financial intake of http://scholar.google.com/scholar_lookup?&title=Use%20of%20a%20multiethnic%20approach%20to%20identify%20rheumatoid-arthritis%E2%80%93susceptibility%20loci%2C%201p36%20and%2017q12&journal=Am.%20J.%20Hum.%20Genet.&doi=10.1016%2Fj.ajhg.2012.01.010&volume=90&pages=524-532&publication_year=2012&author=Kurreeman%2CFA?
• Check the income stats for https://doi.org/10.1038%2Fng.2231
• Check the income stats for http://scholar.google.com/scholar_lookup?&title=Meta-analysis%20identifies%20nine%20new%20loci%20associated%20with%20rheumatoid%20arthritis%20in%20the%20Japanese%20population&journal=Nat.%20Genet.&doi=10.1038%2Fng.2231&volume=44&pages=511-516&publication_year=2012&author=Okada%2CY
• How much cash flow does https://doi.org/10.1016%2FS0140-6736%2812%2960110-X have monthly?
• Check the income stats for http://scholar.google.com/scholar_lookup?&title=The%20interleukin-6%20receptor%20as%20a%20target%20for%20prevention%20of%20coronary%20heart%20disease%3A%20a%20mendelian%20randomisation%20analysis&journal=Lancet&doi=10.1016%2FS0140-6736%2812%2960110-X&volume=379&pages=1214-1224&publication_year=2012&author=Hingorani%2CAD&author=Casas%2CJP
• How much revenue does https://doi.org/10.1016%2FS0140-6736%2811%2961931-4 generate?
• What's the financial gain of http://scholar.google.com/scholar_lookup?&title=Interleukin-6%20receptor%20pathways%20in%20coronary%20heart%20disease%3A%20a%20collaborative%20meta-analysis%20of%2082%20studies&journal=Lancet&doi=10.1016%2FS0140-6736%2811%2961931-4&volume=379&pages=1205-1213&publication_year=2012&author=Sarwar%2CN?
• How much does https://doi.org/10.1016%2FS0140-6736%2811%2960874-X bring in each month?
• Explore the financials of http://scholar.google.com/scholar_lookup?&title=Identification%20of%20IL6R%20and%20chromosome%2011q13.5%20as%20risk%20loci%20for%20asthma&journal=Lancet&doi=10.1016%2FS0140-6736%2811%2960874-X&volume=378&pages=1006-1014&publication_year=2011&author=Ferreira%2CMA
• How much revenue does https://doi.org/10.1002%2Feji.200636400 produce monthly?
• How much profit does http://scholar.google.com/scholar_lookup?&title=Involvement%20of%20GATA3%20in%20protein%20kinase%20C%20%CE%B8%E2%80%93induced%20Th2%20cytokine%20expression&journal=Eur.%20J.%20Immunol.&doi=10.1002%2Feji.200636400&volume=36&pages=3305-3314&publication_year=2006&author=Stevens%2CL generate?
• How profitable is https://doi.org/10.1016%2Fj.ajhg.2011.09.002?
• Learn how profitable http://scholar.google.com/scholar_lookup?&title=Integrating%20autoimmune%20risk%20loci%20with%20gene-expression%20data%20identifies%20specific%20pathogenic%20immune%20cell%20subsets&journal=Am.%20J.%20Hum.%20Genet.&doi=10.1016%2Fj.ajhg.2011.09.002&volume=89&pages=496-506&publication_year=2011&author=Hu%2CX is on a monthly basis
• What's the financial intake of https://doi.org/10.1038%2Fni1008-1091?
• What is the monthly revenue of http://scholar.google.com/scholar_lookup?&title=The%20Immunological%20Genome%20Project%3A%20networks%20of%20gene%20expression%20in%20immune%20cells&journal=Nat.%20Immunol.&doi=10.1038%2Fni1008-1091&volume=9&pages=1091-1094&publication_year=2008&author=Heng%2CTS&author=Painter%2CMW?
• Earnings of https://doi.org/10.1038%2Fnmeth.1785
• How much income is http://scholar.google.com/scholar_lookup?&title=A%20linear%20complexity%20phasing%20method%20for%20thousands%20of%20genomes&journal=Nat.%20Methods&doi=10.1038%2Fnmeth.1785&volume=9&pages=179-181&publication_year=2012&author=Delaneau%2CO&author=Marchini%2CJ&author=Zagury%2CJF earning monthly?
• What's the financial intake of https://doi.org/10.1038%2Fng2088?
• http://scholar.google.com/scholar_lookup?&title=A%20new%20multipoint%20method%20for%20genome-wide%20association%20studies%20by%20imputation%20of%20genotypes&journal=Nat.%20Genet.&doi=10.1038%2Fng2088&volume=39&pages=906-913&publication_year=2007&author=Marchini%2CJ&author=Howie%2CB&author=Myers%2CS&author=McVean%2CG&author=Donnelly%2CP's financial summary
• Financial intake of https://doi.org/10.1086%2F519795
• What's the income generated by http://scholar.google.com/scholar_lookup?&title=PLINK%3A%20a%20tool%20set%20for%20whole-genome%20association%20and%20population-based%20linkage%20analyses&journal=Am.%20J.%20Hum.%20Genet.&doi=10.1086%2F519795&volume=81&pages=559-575&publication_year=2007&author=Purcell%2CS each month?
• Monthly income for https://doi.org/10.1093%2Fhmg%2Fddn288
• What's the financial intake of http://scholar.google.com/scholar_lookup?&title=Practical%20aspects%20of%20imputation-driven%20meta-analysis%20of%20genome-wide%20association%20studies&journal=Hum.%20Mol.%20Genet.&doi=10.1093%2Fhmg%2Fddn288&volume=17&pages=R122-R128&publication_year=2008&author=de%20Bakker%2CPI?
• Find out how much https://doi.org/10.1371%2Fjournal.pgen.1002197 earns monthly
• Find out how much http://scholar.google.com/scholar_lookup?&title=Trans-eQTLs%20reveal%20that%20independent%20genetic%20variants%20associated%20with%20a%20complex%20phenotype%20converge%20on%20intermediate%20genes%2C%20with%20a%20major%20role%20for%20the%20HLA&journal=PLoS%20Genet.&doi=10.1371%2Fjournal.pgen.1002197&volume=7&publication_year=2011&author=Fehrmann%2CRS earns monthly
• Discover the revenue of https://citation-needed.springer.com/v2/references/10.1038/ng.2462?format=refman&flavour=references
• How much revenue does https://www.ncbi.nlm.nih.gov/entrez/query.fcgi?cmd=search&term=Steve%20Eyre generate?
• What is the monthly revenue of https://scholar.google.co.uk/scholar?as_q=&num=10&btnG=Search+Scholar&as_epq=&as_oq=&as_eq=&as_occt=any&as_sauthors=%22Steve%20Eyre%22&as_publication=&as_ylo=&as_yhi=&as_allsubj=all&hl=en?
• https://www.ncbi.nlm.nih.gov/entrez/query.fcgi?cmd=search&term=John%20Bowes's total income per month
• How much profit does https://scholar.google.co.uk/scholar?as_q=&num=10&btnG=Search+Scholar&as_epq=&as_oq=&as_eq=&as_occt=any&as_sauthors=%22John%20Bowes%22&as_publication=&as_ylo=&as_yhi=&as_allsubj=all&hl=en generate?
• What's the monthly money flow for https://www.ncbi.nlm.nih.gov/entrez/query.fcgi?cmd=search&term=Doroth%C3%A9e%20Diogo?
• How much revenue does https://scholar.google.co.uk/scholar?as_q=&num=10&btnG=Search+Scholar&as_epq=&as_oq=&as_eq=&as_occt=any&as_sauthors=%22Doroth%C3%A9e%20Diogo%22&as_publication=&as_ylo=&as_yhi=&as_allsubj=all&hl=en produce monthly?
• What's the income of https://www.ncbi.nlm.nih.gov/entrez/query.fcgi?cmd=search&term=Annette%20Lee?
• Earnings of https://scholar.google.co.uk/scholar?as_q=&num=10&btnG=Search+Scholar&as_epq=&as_oq=&as_eq=&as_occt=any&as_sauthors=%22Annette%20Lee%22&as_publication=&as_ylo=&as_yhi=&as_allsubj=all&hl=en
• How much revenue does https://www.ncbi.nlm.nih.gov/entrez/query.fcgi?cmd=search&term=Anne%20Barton bring in?
• What's the profit of https://scholar.google.co.uk/scholar?as_q=&num=10&btnG=Search+Scholar&as_epq=&as_oq=&as_eq=&as_occt=any&as_sauthors=%22Anne%20Barton%22&as_publication=&as_ylo=&as_yhi=&as_allsubj=all&hl=en?
• How much does https://www.ncbi.nlm.nih.gov/entrez/query.fcgi?cmd=search&term=Paul%20Martin net monthly?
• What is the earnings of https://scholar.google.co.uk/scholar?as_q=&num=10&btnG=Search+Scholar&as_epq=&as_oq=&as_eq=&as_occt=any&as_sauthors=%22Paul%20Martin%22&as_publication=&as_ylo=&as_yhi=&as_allsubj=all&hl=en?
• What's the monthly income of https://www.ncbi.nlm.nih.gov/entrez/query.fcgi?cmd=search&term=Alexandra%20Zhernakova?
• What's the total monthly financial gain of https://scholar.google.co.uk/scholar?as_q=&num=10&btnG=Search+Scholar&as_epq=&as_oq=&as_eq=&as_occt=any&as_sauthors=%22Alexandra%20Zhernakova%22&as_publication=&as_ylo=&as_yhi=&as_allsubj=all&hl=en?
• What's https://www.ncbi.nlm.nih.gov/entrez/query.fcgi?cmd=search&term=Eli%20Stahl's gross income?
• Discover the revenue of https://scholar.google.co.uk/scholar?as_q=&num=10&btnG=Search+Scholar&as_epq=&as_oq=&as_eq=&as_occt=any&as_sauthors=%22Eli%20Stahl%22&as_publication=&as_ylo=&as_yhi=&as_allsubj=all&hl=en
• What are the earnings of https://www.ncbi.nlm.nih.gov/entrez/query.fcgi?cmd=search&term=Sebastien%20Viatte?
• Learn how profitable https://scholar.google.co.uk/scholar?as_q=&num=10&btnG=Search+Scholar&as_epq=&as_oq=&as_eq=&as_occt=any&as_sauthors=%22Sebastien%20Viatte%22&as_publication=&as_ylo=&as_yhi=&as_allsubj=all&hl=en is on a monthly basis
• What's the revenue for https://www.ncbi.nlm.nih.gov/entrez/query.fcgi?cmd=search&term=Kate%20McAllister?
• How much money does https://scholar.google.co.uk/scholar?as_q=&num=10&btnG=Search+Scholar&as_epq=&as_oq=&as_eq=&as_occt=any&as_sauthors=%22Kate%20McAllister%22&as_publication=&as_ylo=&as_yhi=&as_allsubj=all&hl=en make?
• Earnings of https://www.ncbi.nlm.nih.gov/entrez/query.fcgi?cmd=search&term=Christopher%20I%20Amos
• Learn about the earnings of https://scholar.google.co.uk/scholar?as_q=&num=10&btnG=Search+Scholar&as_epq=&as_oq=&as_eq=&as_occt=any&as_sauthors=%22Christopher%20I%20Amos%22&as_publication=&as_ylo=&as_yhi=&as_allsubj=all&hl=en
• Find out how much https://www.ncbi.nlm.nih.gov/entrez/query.fcgi?cmd=search&term=Leonid%20Padyukov earns monthly
• How much does https://scholar.google.co.uk/scholar?as_q=&num=10&btnG=Search+Scholar&as_epq=&as_oq=&as_eq=&as_occt=any&as_sauthors=%22Leonid%20Padyukov%22&as_publication=&as_ylo=&as_yhi=&as_allsubj=all&hl=en gross monthly?
• How much profit does https://www.ncbi.nlm.nih.gov/entrez/query.fcgi?cmd=search&term=Rene%20E%20M%20Toes generate?
• How much revenue does https://scholar.google.co.uk/scholar?as_q=&num=10&btnG=Search+Scholar&as_epq=&as_oq=&as_eq=&as_occt=any&as_sauthors=%22Rene%20E%20M%20Toes%22&as_publication=&as_ylo=&as_yhi=&as_allsubj=all&hl=en produce monthly?
• How much does https://www.ncbi.nlm.nih.gov/entrez/query.fcgi?cmd=search&term=Tom%20W%20J%20Huizinga gross monthly?
• How much profit does https://scholar.google.co.uk/scholar?as_q=&num=10&btnG=Search+Scholar&as_epq=&as_oq=&as_eq=&as_occt=any&as_sauthors=%22Tom%20W%20J%20Huizinga%22&as_publication=&as_ylo=&as_yhi=&as_allsubj=all&hl=en make?
• See how much https://www.ncbi.nlm.nih.gov/entrez/query.fcgi?cmd=search&term=Cisca%20Wijmenga makes per month
• Monthly income for https://scholar.google.co.uk/scholar?as_q=&num=10&btnG=Search+Scholar&as_epq=&as_oq=&as_eq=&as_occt=any&as_sauthors=%22Cisca%20Wijmenga%22&as_publication=&as_ylo=&as_yhi=&as_allsubj=all&hl=en
• Learn how profitable https://www.ncbi.nlm.nih.gov/entrez/query.fcgi?cmd=search&term=Gosia%20Trynka is on a monthly basis
• How much does https://scholar.google.co.uk/scholar?as_q=&num=10&btnG=Search+Scholar&as_epq=&as_oq=&as_eq=&as_occt=any&as_sauthors=%22Gosia%20Trynka%22&as_publication=&as_ylo=&as_yhi=&as_allsubj=all&hl=en bring in each month?
• Explore the financials of https://www.ncbi.nlm.nih.gov/entrez/query.fcgi?cmd=search&term=Lude%20Franke
• Explore the financials of https://scholar.google.co.uk/scholar?as_q=&num=10&btnG=Search+Scholar&as_epq=&as_oq=&as_eq=&as_occt=any&as_sauthors=%22Lude%20Franke%22&as_publication=&as_ylo=&as_yhi=&as_allsubj=all&hl=en
• How much profit is https://www.ncbi.nlm.nih.gov/entrez/query.fcgi?cmd=search&term=Harm-Jan%20Westra making per month?
• What's the income of https://scholar.google.co.uk/scholar?as_q=&num=10&btnG=Search+Scholar&as_epq=&as_oq=&as_eq=&as_occt=any&as_sauthors=%22Harm-Jan%20Westra%22&as_publication=&as_ylo=&as_yhi=&as_allsubj=all&hl=en?
• How much income is https://www.ncbi.nlm.nih.gov/entrez/query.fcgi?cmd=search&term=Lars%20Alfredsson earning monthly?
• Get to know what's the income of https://scholar.google.co.uk/scholar?as_q=&num=10&btnG=Search+Scholar&as_epq=&as_oq=&as_eq=&as_occt=any&as_sauthors=%22Lars%20Alfredsson%22&as_publication=&as_ylo=&as_yhi=&as_allsubj=all&hl=en
• What's the total monthly financial gain of https://www.ncbi.nlm.nih.gov/entrez/query.fcgi?cmd=search&term=Xinli%20Hu?
• What's the financial outcome of https://scholar.google.co.uk/scholar?as_q=&num=10&btnG=Search+Scholar&as_epq=&as_oq=&as_eq=&as_occt=any&as_sauthors=%22Xinli%20Hu%22&as_publication=&as_ylo=&as_yhi=&as_allsubj=all&hl=en?
• What are the total earnings of https://www.ncbi.nlm.nih.gov/entrez/query.fcgi?cmd=search&term=Cynthia%20Sandor?
• How much does https://scholar.google.co.uk/scholar?as_q=&num=10&btnG=Search+Scholar&as_epq=&as_oq=&as_eq=&as_occt=any&as_sauthors=%22Cynthia%20Sandor%22&as_publication=&as_ylo=&as_yhi=&as_allsubj=all&hl=en make?
• How much does https://www.ncbi.nlm.nih.gov/entrez/query.fcgi?cmd=search&term=Paul%20I%20W%20de%20Bakker earn?
• How much money does https://scholar.google.co.uk/scholar?as_q=&num=10&btnG=Search+Scholar&as_epq=&as_oq=&as_eq=&as_occt=any&as_sauthors=%22Paul%20I%20W%20de%20Bakker%22&as_publication=&as_ylo=&as_yhi=&as_allsubj=all&hl=en generate?
• What are the total earnings of https://www.ncbi.nlm.nih.gov/entrez/query.fcgi?cmd=search&term=Sonia%20Davila?
• How much profit is https://scholar.google.co.uk/scholar?as_q=&num=10&btnG=Search+Scholar&as_epq=&as_oq=&as_eq=&as_occt=any&as_sauthors=%22Sonia%20Davila%22&as_publication=&as_ylo=&as_yhi=&as_allsubj=all&hl=en making per month?
• How much profit does https://www.ncbi.nlm.nih.gov/entrez/query.fcgi?cmd=search&term=Chiea%20Chuen%20Khor make?
• What's the financial outcome of https://scholar.google.co.uk/scholar?as_q=&num=10&btnG=Search+Scholar&as_epq=&as_oq=&as_eq=&as_occt=any&as_sauthors=%22Chiea%20Chuen%20Khor%22&as_publication=&as_ylo=&as_yhi=&as_allsubj=all&hl=en?
• What is the earnings of https://www.ncbi.nlm.nih.gov/entrez/query.fcgi?cmd=search&term=Khai%20Koon%20Heng?
• Financial intake of https://scholar.google.co.uk/scholar?as_q=&num=10&btnG=Search+Scholar&as_epq=&as_oq=&as_eq=&as_occt=any&as_sauthors=%22Khai%20Koon%20Heng%22&as_publication=&as_ylo=&as_yhi=&as_allsubj=all&hl=en
• How much money does https://www.ncbi.nlm.nih.gov/entrez/query.fcgi?cmd=search&term=Robert%20Andrews make?
• See how much https://scholar.google.co.uk/scholar?as_q=&num=10&btnG=Search+Scholar&as_epq=&as_oq=&as_eq=&as_occt=any&as_sauthors=%22Robert%20Andrews%22&as_publication=&as_ylo=&as_yhi=&as_allsubj=all&hl=en makes per month
• https://www.ncbi.nlm.nih.gov/entrez/query.fcgi?cmd=search&term=Sarah%20Edkins's total income per month
• How much does https://scholar.google.co.uk/scholar?as_q=&num=10&btnG=Search+Scholar&as_epq=&as_oq=&as_eq=&as_occt=any&as_sauthors=%22Sarah%20Edkins%22&as_publication=&as_ylo=&as_yhi=&as_allsubj=all&hl=en generate monthly?
• What's the total monthly financial gain of https://www.ncbi.nlm.nih.gov/entrez/query.fcgi?cmd=search&term=Sarah%20E%20Hunt?
• Income figures for https://scholar.google.co.uk/scholar?as_q=&num=10&btnG=Search+Scholar&as_epq=&as_oq=&as_eq=&as_occt=any&as_sauthors=%22Sarah%20E%20Hunt%22&as_publication=&as_ylo=&as_yhi=&as_allsubj=all&hl=en
• Earnings of https://www.ncbi.nlm.nih.gov/entrez/query.fcgi?cmd=search&term=Cordelia%20Langford
• How much income does https://scholar.google.co.uk/scholar?as_q=&num=10&btnG=Search+Scholar&as_epq=&as_oq=&as_eq=&as_occt=any&as_sauthors=%22Cordelia%20Langford%22&as_publication=&as_ylo=&as_yhi=&as_allsubj=all&hl=en have?
• Revenue of https://www.ncbi.nlm.nih.gov/entrez/query.fcgi?cmd=search&term=Deborah%20Symmons
• What is the monthly revenue of https://scholar.google.co.uk/scholar?as_q=&num=10&btnG=Search+Scholar&as_epq=&as_oq=&as_eq=&as_occt=any&as_sauthors=%22Deborah%20Symmons%22&as_publication=&as_ylo=&as_yhi=&as_allsubj=all&hl=en?
• How much profit does https://www.ncbi.nlm.nih.gov/entrez/query.fcgi?cmd=search&term=Pat%20Concannon make?
• What is the earnings of https://scholar.google.co.uk/scholar?as_q=&num=10&btnG=Search+Scholar&as_epq=&as_oq=&as_eq=&as_occt=any&as_sauthors=%22Pat%20Concannon%22&as_publication=&as_ylo=&as_yhi=&as_allsubj=all&hl=en?
• Earnings of https://www.ncbi.nlm.nih.gov/entrez/query.fcgi?cmd=search&term=Suna%20Onengut-Gumuscu
• Get to know https://scholar.google.co.uk/scholar?as_q=&num=10&btnG=Search+Scholar&as_epq=&as_oq=&as_eq=&as_occt=any&as_sauthors=%22Suna%20Onengut-Gumuscu%22&as_publication=&as_ylo=&as_yhi=&as_allsubj=all&hl=en's earnings
• Earnings of https://www.ncbi.nlm.nih.gov/entrez/query.fcgi?cmd=search&term=Stephen%20S%20Rich
• How much does https://scholar.google.co.uk/scholar?as_q=&num=10&btnG=Search+Scholar&as_epq=&as_oq=&as_eq=&as_occt=any&as_sauthors=%22Stephen%20S%20Rich%22&as_publication=&as_ylo=&as_yhi=&as_allsubj=all&hl=en gross monthly?
• How much does https://www.ncbi.nlm.nih.gov/entrez/query.fcgi?cmd=search&term=Panos%20Deloukas net monthly?
• How much revenue does https://scholar.google.co.uk/scholar?as_q=&num=10&btnG=Search+Scholar&as_epq=&as_oq=&as_eq=&as_occt=any&as_sauthors=%22Panos%20Deloukas%22&as_publication=&as_ylo=&as_yhi=&as_allsubj=all&hl=en produce monthly?
• Find out how much https://www.ncbi.nlm.nih.gov/entrez/query.fcgi?cmd=search&term=Miguel%20A%20Gonzalez-Gay earns monthly
• What's the financial intake of https://scholar.google.co.uk/scholar?as_q=&num=10&btnG=Search+Scholar&as_epq=&as_oq=&as_eq=&as_occt=any&as_sauthors=%22Miguel%20A%20Gonzalez-Gay%22&as_publication=&as_ylo=&as_yhi=&as_allsubj=all&hl=en?
• How much income does https://www.ncbi.nlm.nih.gov/entrez/query.fcgi?cmd=search&term=Luis%20Rodriguez-Rodriguez have?
• How much does https://scholar.google.co.uk/scholar?as_q=&num=10&btnG=Search+Scholar&as_epq=&as_oq=&as_eq=&as_occt=any&as_sauthors=%22Luis%20Rodriguez-Rodriguez%22&as_publication=&as_ylo=&as_yhi=&as_allsubj=all&hl=en pull in?
• Revenue of https://www.ncbi.nlm.nih.gov/entrez/query.fcgi?cmd=search&term=Lisbeth%20%C3%84rlsetig
• What's the financial outcome of https://scholar.google.co.uk/scholar?as_q=&num=10&btnG=Search+Scholar&as_epq=&as_oq=&as_eq=&as_occt=any&as_sauthors=%22Lisbeth%20%C3%84rlsetig%22&as_publication=&as_ylo=&as_yhi=&as_allsubj=all&hl=en?
• Check the income stats for https://www.ncbi.nlm.nih.gov/entrez/query.fcgi?cmd=search&term=Javier%20Martin
• How much profit does https://scholar.google.co.uk/scholar?as_q=&num=10&btnG=Search+Scholar&as_epq=&as_oq=&as_eq=&as_occt=any&as_sauthors=%22Javier%20Martin%22&as_publication=&as_ylo=&as_yhi=&as_allsubj=all&hl=en generate?
• What's the revenue for https://www.ncbi.nlm.nih.gov/entrez/query.fcgi?cmd=search&term=Solbritt%20Rantap%C3%A4%C3%A4-Dahlqvist?
• Income figures for https://scholar.google.co.uk/scholar?as_q=&num=10&btnG=Search+Scholar&as_epq=&as_oq=&as_eq=&as_occt=any&as_sauthors=%22Solbritt%20Rantap%C3%A4%C3%A4-Dahlqvist%22&as_publication=&as_ylo=&as_yhi=&as_allsubj=all&hl=en
• How much revenue does https://www.ncbi.nlm.nih.gov/entrez/query.fcgi?cmd=search&term=Robert%20M%20Plenge produce monthly?
• Explore the financials of https://scholar.google.co.uk/scholar?as_q=&num=10&btnG=Search+Scholar&as_epq=&as_oq=&as_eq=&as_occt=any&as_sauthors=%22Robert%20M%20Plenge%22&as_publication=&as_ylo=&as_yhi=&as_allsubj=all&hl=en
• https://www.ncbi.nlm.nih.gov/entrez/query.fcgi?cmd=search&term=Soumya%20Raychaudhuri's total income per month
• How much does https://scholar.google.co.uk/scholar?as_q=&num=10&btnG=Search+Scholar&as_epq=&as_oq=&as_eq=&as_occt=any&as_sauthors=%22Soumya%20Raychaudhuri%22&as_publication=&as_ylo=&as_yhi=&as_allsubj=all&hl=en earn?
• See how much https://www.ncbi.nlm.nih.gov/entrez/query.fcgi?cmd=search&term=Lars%20Klareskog makes per month
• How much revenue does https://scholar.google.co.uk/scholar?as_q=&num=10&btnG=Search+Scholar&as_epq=&as_oq=&as_eq=&as_occt=any&as_sauthors=%22Lars%20Klareskog%22&as_publication=&as_ylo=&as_yhi=&as_allsubj=all&hl=en generate?
• What is the monthly revenue of https://www.ncbi.nlm.nih.gov/entrez/query.fcgi?cmd=search&term=Peter%20K%20Gregersen?
• How much does https://scholar.google.co.uk/scholar?as_q=&num=10&btnG=Search+Scholar&as_epq=&as_oq=&as_eq=&as_occt=any&as_sauthors=%22Peter%20K%20Gregersen%22&as_publication=&as_ylo=&as_yhi=&as_allsubj=all&hl=en make?
• How much does https://www.ncbi.nlm.nih.gov/entrez/query.fcgi?cmd=search&term=Jane%20Worthington bring in each month?
• How much does https://scholar.google.co.uk/scholar?as_q=&num=10&btnG=Search+Scholar&as_epq=&as_oq=&as_eq=&as_occt=any&as_sauthors=%22Jane%20Worthington%22&as_publication=&as_ylo=&as_yhi=&as_allsubj=all&hl=en earn?
• Financial intake of https://static-content.springer.com/esm/art%3A10.1038%2Fng.2462/MediaObjects/41588_2012_BFng2462_MOESM35_ESM.pdf
• Learn about the earnings of https://static-content.springer.com/esm/art%3A10.1038%2Fng.2462/MediaObjects/41588_2012_BFng2462_MOESM36_ESM.xls
• Learn about the earnings of https://s100.copyright.com/AppDispatchServlet?title=High-density%20genetic%20mapping%20identifies%20new%20susceptibility%20loci%20for%20rheumatoid%20arthritis&author=Steve%20Eyre%20et%20al&contentID=10.1038%2Fng.2462©right=Springer%20Nature%20America%2C%20Inc.&publication=1061-4036&publicationDate=2012-11-11&publisherName=SpringerNature&orderBeanReset=true
• https://citation-needed.springer.com/v2/references/10.1038/ng.2462?format=refman&flavour=citation income
• How much money does https://doi.org/10.1186/s13075-023-03245-x make?
• How much does https://doi.org/10.1038/s41431-022-01264-x net monthly?
• How much profit does https://doi.org/10.1038/s41588-024-01732-8 generate?
• https://doi.org/10.1186/s12864-023-09227-2 income
• Learn how profitable https://doi.org/10.1186/s12863-022-01102-5 is on a monthly basis
• How much profit is https://www.protocols.io/ making per month?
• How much money does https://www.natureindex.com/ make?
• How much does http://www.naturechina.com generate monthly?
• How much income is https://www.natureasia.com/ja-jp earning monthly?

Analytics and Tracking {📊}

• Google Tag Manager

Libraries {📚}

• Prism.js
• Zoom.js

Emails and Hosting {✉️}

Mail Servers:

• mxa-002c5801.gslb.pphosted.com
• mxb-002c5801.gslb.pphosted.com

Name Servers:

• pdns1.ultradns.net
• pdns2.ultradns.net
• pdns3.ultradns.org
• pdns4.ultradns.org
• pdns5.ultradns.info
• pdns6.ultradns.co.uk

CDN Services {📦}

• Crossref